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A fen is a type of
peat Peat (), also known as turf (), is an accumulation of partially decayed vegetation or organic matter. It is unique to natural areas called peatlands, bogs, mires, moors, or muskegs. The peatland ecosystem covers and is the most efficien ...
-accumulating
wetland A wetland is a distinct ecosystem that is flooded or saturated by water, either permanently (for years or decades) or seasonally (for weeks or months). Flooding results in oxygen-free (anoxic) processes prevailing, especially in the soils. The ...
fed by mineral-rich ground or
surface water Surface water is water located on top of land forming terrestrial (inland) waterbodies, and may also be referred to as ''blue water'', opposed to the seawater and waterbodies like the ocean. The vast majority of surface water is produced by prec ...
. It is one of the main types of wetlands along with
marsh A marsh is a wetland that is dominated by herbaceous rather than woody plant species.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p Marshes can often be found at ...
es,
swamp A swamp is a forested wetland.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p. Swamps are considered to be transition zones because both land and water play a role in ...
s, and bogs. Bogs and fens, both peat-forming ecosystems, are also known as
mire A mire, peatland, or quagmire is a wetland area dominated by living peat-forming plants. Mires arise because of incomplete decomposition of organic matter, usually litter from vegetation, due to water-logging and subsequent anoxia. All types ...
s. The unique
water chemistry Water chemistry analyses are carried out to identify and quantify the chemical components and properties of water samples. The type and sensitivity of the analysis depends on the purpose of the analysis and the anticipated use of the water. Chemic ...
of fens is a result of the ground or surface water input. Typically, this input results in higher mineral concentrations and a more
basic BASIC (Beginners' All-purpose Symbolic Instruction Code) is a family of general-purpose, high-level programming languages designed for ease of use. The original version was created by John G. Kemeny and Thomas E. Kurtz at Dartmouth College ...
pH than found in bogs. As peat accumulates in a fen,
groundwater Groundwater is the water present beneath Earth's surface in rock and soil pore spaces and in the fractures of rock formations. About 30 percent of all readily available freshwater in the world is groundwater. A unit of rock or an unconsolidate ...
input can be reduced or cut off, making the fen
ombrotrophic Ombrotrophic ("cloud-fed"), from Ancient Greek ὄμβρος (''ómvros'') meaning "rain" and τροφή (''trofí'') meaning "food"), refers to Soil, soils or vegetation which receive all of their water and nutrients from precipitation, rather ...
rather than
minerotrophic Minerotrophic refers to environments that receive nutrients primarily through groundwater that flows through mineral-rich soils or rock,Environment Canada (2014). Ontario wetland evaluation system: Northern Manual, 1st edition, version 3.2. Queen’ ...
. In this way, fens can become more acidic and transition to bogs over time. Fens can be found around the world, but the vast majority are located at the mid to high latitudes of the Northern Hemisphere. They are dominated by
sedges The Cyperaceae are a family of graminoid (grass-like), monocotyledonous flowering plants known as sedges. The family is large, with some 5,500 known species described in about 90 genera, the largest being the "true sedges" genus ''Carex'' wit ...
and
moss Mosses are small, non-vascular flowerless plants in the taxonomic division Bryophyta (, ) '' sensu stricto''. Bryophyta (''sensu lato'', Schimp. 1879) may also refer to the parent group bryophytes, which comprise liverworts, mosses, and hor ...
es, particularly
graminoid In botany and ecology, graminoid refers to a herbaceous plant with a grass-like morphology, i.e. elongated culms with long, blade-like leaves. They are contrasted to forbs, herbaceous plants without grass-like features. The plants most ofte ...
s that may be rarely found elsewhere, such as the sedge species ''
Carex exilis ''Carex exilis'', common name coastal sedge or meager sedge, is a species of ''Carex'' native to North America, with several disjunct populations from southern Canada to the Gulf Coast. It is rare in much of its range and is listed as an endanger ...
''. Fens are highly biodiverse ecosystems and often serve as habitats for endangered or rare species, with species composition changing with water chemistry. They also play important roles in the cycling of nutrients such as carbon, nitrogen, and
phosphorus Phosphorus is a chemical element with the symbol P and atomic number 15. Elemental phosphorus exists in two major forms, white phosphorus and red phosphorus, but because it is highly reactive, phosphorus is never found as a free element on Ear ...
due to the lack of oxygen (anaerobic conditions) in waterlogged organic fen soils. Fens have historically been converted to agricultural land. However, fens face a number of other threats, including peat cutting, pollution, invasive species, and nearby disturbances that lower the water table in the fen, such as quarrying. Interrupting the flow of mineral-rich water into a fen changes the water chemistry, which can alter
species richness Species richness is the number of different species represented in an ecological community, landscape or region. Species richness is simply a count of species, and it does not take into account the abundances of the species or their relative a ...
and dry out the peat. Drier peat is more easily decomposed and can even burn.


Distribution and extent

Fens are distributed around the world, but are most frequently found at the mid-high latitudes of the Northern Hemisphere. They are found throughout the
temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout t ...
zone and
boreal Boreal may refer to: Climatology and geography *Boreal (age), the first climatic phase of the Blytt-Sernander sequence of northern Europe, during the Holocene epoch *Boreal climate, a climate characterized by long winters and short, cool to mild ...
regions, but are also present in
tundra In physical geography, tundra () is a type of biome where tree growth is hindered by frigid temperatures and short growing seasons. The term ''tundra'' comes through Russian (') from the Kildin Sámi word (') meaning "uplands", "treeless moun ...
and in specific environmental conditions in other regions around the world. In the United States, fens are most common in the Midwest and Northeast, but can be found across the country. In Canada, fens are most frequent in the
lowlands Upland and lowland are conditional descriptions of a plain based on elevation above sea level. In studies of the ecology of freshwater rivers, habitats are classified as upland or lowland. Definitions Upland and lowland are portions of p ...
near
Hudson Bay Hudson Bay ( crj, text=ᐐᓂᐯᒄ, translit=Wînipekw; crl, text=ᐐᓂᐹᒄ, translit=Wînipâkw; iu, text=ᑲᖏᖅᓱᐊᓗᒃ ᐃᓗᐊ, translit=Kangiqsualuk ilua or iu, text=ᑕᓯᐅᔭᕐᔪᐊᖅ, translit=Tasiujarjuaq; french: b ...
and
James Bay James Bay (french: Baie James; cr, ᐐᓂᐯᒄ, Wînipekw, dirty water) is a large body of water located on the southern end of Hudson Bay in Canada. Both bodies of water extend from the Arctic Ocean, of which James Bay is the southernmost par ...
, but can also be found across the country. Fens are also spread across the northern latitudes of Eurasia, including Britain and Ireland, as well as Japan, but east-central Europe is especially rich in fens. Further south, fens are much rarer, but do exist under specific conditions. In Africa, fens have been found in the
Okavango Delta The Okavango Delta (or Okavango Grassland; formerly spelled "Okovango" or "Okovanggo") in Botswana is a swampy inland delta formed where the Okavango River reaches a tectonic trough at an altitude of 930–1,000 m in the central part of the en ...
in Botswana and the highland slopes in
Lesotho Lesotho ( ), officially the Kingdom of Lesotho, is a country landlocked country, landlocked as an Enclave and exclave, enclave in South Africa. It is situated in the Maloti Mountains and contains the Thabana Ntlenyana, highest mountains in Sou ...
. Fens can also be found at the colder latitudes of the Southern Hemisphere. They are found in New Zealand and southwest Argentina, but the extent is much less than that of the northern latitudes. Locally, fens are most often found at the intersection of terrestrial and aquatic ecosystems, such as the headwaters of streams and rivers. It is estimated that there are approximately 1.1 million square kilometers of fens worldwide, but quantifying the extent of fens is difficult. Because wetland definitions vary regionally, not all countries define fens the same way. In addition, wetland data is not always available or of high quality. Fens are also difficult to rigidly delineate and measure, as they are located between terrestrial and aquatic ecosystems.


Definition

Rigidly defining types of wetlands, including fens, is difficult for a number of reasons. First, wetlands are diverse and varied ecosystems that are not easily categorized according to inflexible definitions. They are often described as a transition between
terrestrial Terrestrial refers to things related to land or the planet Earth. Terrestrial may also refer to: * Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on or near the ground, as opposed to ...
and aquatic ecosystems with characteristics of both. This makes it difficult to delineate the exact extent of a wetland. Second, terms used to describe wetland types vary greatly by region. The term ''
bayou In usage in the Southern United States, a bayou () is a body of water typically found in a flat, low-lying area. It may refer to an extremely slow-moving stream, river (often with a poorly defined shoreline), marshy lake, wetland, or creek. They ...
,'' for example, describes a type of wetland, but its use is generally limited to the southern United States. Third, different languages use different terms to describe types of wetlands. For instance, in Russian, there is no equivalent word for the term ''
swamp A swamp is a forested wetland.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p. Swamps are considered to be transition zones because both land and water play a role in ...
'' as it is typically used in North America. The result is a large number of
wetland classification Classification of wetlands has been a problematical task, with the commonly accepted definition of what constitutes a wetland being among the major difficulties. A number of national wetland classifications exist. In the 1970s, the Ramsar Conventio ...
systems that each define wetlands and wetland types in their own way. However, many classification systems include four broad categories that most wetlands fall into:
marsh A marsh is a wetland that is dominated by herbaceous rather than woody plant species.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p Marshes can often be found at ...
, swamp, bog, and fen. While classification systems differ on the exact criteria that define a fen, there are common characteristics that describe fens generally and imprecisely. A general definition provided by the textbook ''Wetlands'' describes a fen as "a peat-accumulating wetland that receives some drainage from surrounding mineral soil and usually supports marsh like vegetation." Three examples are presented below to illustrate more specific definitions for the term ''fen''.


Canadian Wetland Classification System definition

In the Canadian Wetland Classification System, fens are defined by six characteristics: # Peat is present. # The surface of the wetland is level with the
water table The water table is the upper surface of the zone of saturation. The zone of saturation is where the pores and fractures of the ground are saturated with water. It can also be simply explained as the depth below which the ground is saturated. T ...
. Water flows on the surface and through the subsurface of the wetland. # The water table fluctuates. It may be at the surface of the wetland or a few centimeters above or below it. # The wetland receives a significant amount of its water from mineral-rich groundwater or surface water. # Decomposed
sedges The Cyperaceae are a family of graminoid (grass-like), monocotyledonous flowering plants known as sedges. The family is large, with some 5,500 known species described in about 90 genera, the largest being the "true sedges" genus ''Carex'' wit ...
or brown moss peat are present. # The vegetation is predominantly