Felicia Nordenstamii
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''Felicia nordenstamii'' is a flowering shrub in the family Asteraceae. It is found only in South Africa where it grows on limestone hills close to the sea on the southern coast. ''Felicia nordenstamii'' is a many-branched shrub growing up to tall. The lower parts of the stems are covered in grayish brown bark and the upper stem has many crowded, upwardly angled, alternate leaves with long hairs on the lower surfaces. Large flower heads form at the tips of the branches, each about 4 cm (1 in) across, with about thirty purplish blue ray florets surrounding many yellow disc florets.


Description

''Felicia nordenstamii'' is a branched
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
of up to high, that is relatively strongly woody at its base. Where it has lost its leaves in lower part of the stem, it is covered with grayish brown bark. In the leaf-covered younger parts, the stem carries long hairs. The leaves are set alternately along the stem, densely crowded, at an upward angle, without a
stalk Stalk or stalking may refer to: Behaviour * Stalk, the stealthy approach (phase) of a predator towards its prey * Stalking, an act of intrusive behaviour or unwanted attention towards a person * Deer stalking, the pursuit of deer for sport Biol ...
, narrowly inverted egg-shaped, about long and wide, with an entire margin that is somewhat curved upwards, the lower surface with long hairs, and the upper surface hairless. The large
flower heads A pseudanthium (Greek for "false flower"; ) is an inflorescence that resembles a flower. The word is sometimes used for other structures that are neither a true flower nor a true inflorescence. Examples of pseudanthia include flower heads, compos ...
are set individually at the end of flower stalks of up to 1 cm (0.4 in) long, covered in dense long woolly hair. Surrounding each flower head are three to four rows of unruly arranged
bract In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves. They may be smaller, larger, or of ...
s (or phyllaries) that together form the so-called involucre, which is up to in diameter. These bracts are lance-shaped, about wide, and of varying length. The outer bracts are long-haired, about long, the inner eventually hairless, about long. Approximately thirty female ray florets, have purplish blue straps of about 15 mm (0.6 in) long and 2 mm (0.08 in) wide, and are hairy in the upper part of the tubular section at its base. These surround numerous bisexual disc florets with a yellow
corolla Corolla may refer to: *Corolla (botany), the petals of a flower, considered as a unit *Toyota Corolla, an automobile model name * Corolla (headgear), an ancient headdress in the form of a circlet or crown * ''Corolla'' (gastropod), a genus of moll ...
of about long, hairy in the middle part. In the center of each
corolla Corolla may refer to: *Corolla (botany), the petals of a flower, considered as a unit *Toyota Corolla, an automobile model name * Corolla (headgear), an ancient headdress in the form of a circlet or crown * ''Corolla'' (gastropod), a genus of moll ...
are five
anthers The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filam ...
merged into a tube, through which the style grows when the floret opens, hoovering up the
pollen Pollen is a powdery substance produced by seed plants. It consists of pollen grains (highly reduced microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametophyt ...
on its shaft. At the tip of both style branches is a narrowly triangular appendage. Surrounding the base of the corolla are many, yellowish white, serrated, more or less deciduous pappus bristles, all about equal in length at . The eventually yellowish brown to reddish, dry, one-seeded, indehiscent fruits called cypselae are elliptic in outline, about long and wide, with a ridge along the margin. The cypselae of the ray florets are hairless, those of the disc florets short-haired. Flowering occurs from September to October.


Differences with related species

Like '' Felicia echinata'' and '' F. westae'', together with which ''F. nordenstamii'' constitutes the section '' Anhebecarpaea'', it has firm, very regularly and densely set leaves on the younger parts of the stems, distinguishing them from all other ''Felicia'' species. The dense, long and woolly hairs, the long inner involucral bracts, and the narrowly obovate leaves set at an upward angle, set it apart from ''F. echinata'' (recurved narrowly ovate leaves) and ''F. westae'' (line- to lance-shaped leaves, pressed against the stem, hairless except for a fringe). It also has a likeness to ''
Polyarrhena imbricata ''Polyarrhena'' is a genus of low, branching shrublets that is assigned to the Asteraceae, daisy family (Compositae or Asteraceae). Its stems are alternately and densely set with entire or somewhat toothed leaves. Like in almost all Asteraceae, t ...
'', which lacks the woolly hairs, large involucral bracts, and has white ray florets, tinged purple on the underside.


Taxonomy

As far as known, ''Felicia nordenstamii'' was first collected just east of Cape Agulhas by Terence Macleane Salter in 1934. It was recognized as a distinct species by in his 1973 ''
Revision Revision is the process of revising. More specifically, it may refer to: * Patch (computing), Update, a modification of software or a database * Revision control, the management of changes to sets of computer files * ''ReVisions'', a 2004 antholo ...
of the genus Felicia (Asteraceae)''. The species is considered to be part of the section Anhebecarpaea. The species was named in honor of
Rune Bertil Nordenstam Rune Bertil Nordenstam (born 1936) is a Sweden, Swedish botanist and professor emeritus at the Swedish Museum of Natural History in the Department of Phanerogamic Botany. He has worked with Colchicaceae, Senecioneae and Calenduleae, was the edito ...
, a Swedish botanist that collected two specimens of this plant in 1962.


Distribution, habitat and ecology

''Felicia nordenstamii'' is only known from limestone ridges just east of Cape Agulhas and near the Potberg.


Conservation

''Felicia nordenstamii'' is considered near threatened. It grows only at about fifteen locations spread over and area of about . A substantial part of its habitat was lost over due to urban expansion around Struisbaai. It grows on several limestone ridges that have been identified for development. Its habitat is also degraded over its entire range because of invasive ''Acacia'' species.


References


External links


Photos of ''Felicia nordenstamii''
on iNaturalist
Line drawing of ''Felicia nordenstamii''

Distribution of ''Felicia nordenstamii''
{{Taxonbar, from=Q15598252 nordenstamii Endemic flora of the Cape Provinces Plants described in 1973