Feilongus Youngi
   HOME

TheInfoList



OR:

''Feilongus'' is an
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
ctenochasmatid Ctenochasmatidae is a group of pterosaurs within the suborder Pterodactyloidea. They are characterized by their distinctive teeth, which are thought to have been used for filter-feeding. Ctenochasmatids lived from the Late Jurassic to the Early C ...
pterodactyloid Pterodactyloidea (derived from the Greek words ''πτερόν'' (''pterón'', for usual ''ptéryx'') "wing", and ''δάκτυλος'' (''dáktylos'') "finger" meaning "winged finger", "wing-finger" or "finger-wing") is one of the two traditional ...
pterosaur Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to ...
from the
Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 129.4 ± 1.5 Ma (million years ago) and 121.4 ± 1.0 Ma). It is a subdivision of the Early Cretaceous Epoch (or Lower Cretaceous Series). It is preceded ...
Aptian The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ago), a ...
-age
Lower Cretaceous Lower may refer to: *Lower (surname) *Lower Township, New Jersey *Lower Receiver (firearms) *Lower Wick Lower Wick is a small hamlet located in the county of Gloucestershire, England. It is situated about five miles south west of Dursley, eight ...
Yixian Formation The Yixian Formation (; formerly transcribed as Yihsien Formation) is a geological formation in Jinzhou, Liaoning, People's Republic of China, that spans the late Barremian and early Aptian stages of the Early Cretaceous. It is known for its ex ...
of
Beipiao Beipiao () is a city in Chaoyang prefecture, Liaoning province, in Northeast China. It has a population of 202,807. The main industry in the area is coal mining. With vertical shafts of almost 1000m, these are some of the deepest coal mines in C ...
, Liaoning, China.


Discovery and naming

The genus was named and described in 2005 by
Wang Xiaolin Wang may refer to: Names * Wang (surname) (王), a common Chinese surname * Wāng (汪), a less common Chinese surname * Titles in Chinese nobility * A title in Korean nobility * A title in Mongolian nobility Places * Wang River in Thailand ...
,
Alexander Kellner Alexander Wilhelm Armin Kellner (born September 26, 1961) is a Brazilian geologist and paleontologist who is a leading expert in the field of studying pterosaurs. His research has focused mainly on fossil reptiles from the Cretaceous Period, i ...
,
Zhou Zhonghe Zhou Zhonghe (; born 19 January 1965 in Jiangdu, Jiangsu province) is a Chinese palaeontologist. He described the ancient bird ''Confuciusornis''.. Zhou graduated from Nanjing University and earned a Ph.D. in Biology in 1999 from the University o ...
and
Diogenes de Almeida Campos Diogenes ( ; grc, Διογένης, Diogénēs ), also known as Diogenes the Cynic (, ) or Diogenes of Sinope, was a Greek philosopher and one of the founders of Cynicism (philosophy). He was born in Sinope, an Ionian colony on the Black Sea ...
. The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
is ''Feilongus youngi''. The genus name is derived from ''
Feilong ''Feilong'' (; lit. "flying dragon") is a legendary creature that flies among clouds in Chinese mythology. Feilong is a proper name, and is often used as a title for other ideas and objects. Word The Chinese dragon name ''feilong'' combines ''fei ...
'', the "flying dragon". The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
honors the Chinese
paleontologist Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fossi ...
Yang Zhongjian Yang Zhongjian, also Yang Chung-chien (; 1 June 1897 – 15 January 1979), courtesy name Keqiang (), also known as C.C. (Chung Chien) Young, was a Chinese paleontologist and zoologist. He was one of China's foremost vertebrate paleontologists. H ...
(C. C. Young). ''Feilongus'' is based on
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
IVPP V-12539, a skull and articulated mandible, with on the same plate the detached posterior braincase, of a subadult individual. The fossil is strongly crushed. In 2014, a second specimen, DNMHM D3068 found at Gonggao, was referred to a ''Feilongus'' sp. It consists of a skull with lower jaws and four neck vertebrae. It was a possible subadult or, despite a smaller size, adult.


Description

The
wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingspan of ...
of ''Feilongus'' was estimated by Wang and colleagues to have been around 2.4 meters (7.9 feet), making it large for a basal pterodactyloid. ''Feilongus'' is notable for having two bony crests on the skull (one long and low on middle of the
snout A snout is the protruding portion of an animal's face, consisting of its nose, mouth, and jaw. In many animals, the structure is called a muzzle, rostrum, or proboscis. The wet furless surface around the nostrils of the nose of many mammals is c ...
, and one projecting backwards from the rear of the skull), and for the upper jaws being 10% or longer than the lower jaws, giving it a pronounced
overbite Overbite is the extent of vertical ( superior-inferior) overlap of the maxillary central incisors over the mandibular central incisors, measured relative to the incisal ridges. The term overbite does not refer to a specific condition, nor is ...
. The second specimen however, shows neither crests nor overbite. The preserved part of the second crest was short with the leading edge rounded, and may have had a nonbony extension, now lost. The skull of the holotype is 390–400 millimeters long (15.4–15.7 inches) and extremely elongated with a slightly concave top. The skull and lower jaws held 76 long, curved needle-like
teeth A tooth ( : teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tear ...
, eighteen in the upper, nineteen in the lower jaw, confined to the beak ends, the anterior third, of the jaws. The second specimen had seventy-eight teeth. The neck vertebrae of the second specimen are very elongated, five times longer than wide.


Phylogeny

A
cladistic Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived char ...
analysis by the describers showed ''Feilongus'' as the sister taxon of a
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
consisting of ''
Gallodactylus ''Cycnorhamphus'' (meaning "swan beak") is a genus of gallodactylid ctenochasmatoid pterosaur from the Late Jurassic period of France and Germany, about 152 million years ago. It is probably synonymous with the genus ''Gallodactylus''. History ...
'' and ''
Cycnorhamphus ''Cycnorhamphus'' (meaning "swan beak") is a genus of gallodactylid ctenochasmatoid pterosaur from the Late Jurassic period of France and Germany, about 152 million years ago. It is probably synonymous with the genus ''Gallodactylus''. History ...
'', meaning it was a member of the
Gallodactylidae Gallodactylidae is a group of pterosaurs within the suborder Pterodactyloidea. Gallodactylids differed from other related pterosaurs in several distinct features, including fewer than 50 teeth present only in the jaw tips, and rounded crests pres ...
''sensu'' Kellner, a group of ctenochasmatoids, within the larger
Archaeopterodactyloidea Archaeopterodactyloidea (meaning "ancient Pterodactyloidea") is an extinct clade of pterodactyloid pterosaurs that lived from the middle Late Jurassic to the latest Early Cretaceous periods (Kimmeridgian to Albian stages) of Africa, Asia, Europe ...
, the clade containing according to
Alexander Kellner Alexander Wilhelm Armin Kellner (born September 26, 1961) is a Brazilian geologist and paleontologist who is a leading expert in the field of studying pterosaurs. His research has focused mainly on fossil reptiles from the Cretaceous Period, i ...
the most basal pterodactyloids. The Ctenochasmatoidea are known for having numerous small, thin teeth, possibly for straining food from water, as
flamingo Flamingos or flamingoes are a type of Wader, wading bird in the Family (biology), family Phoenicopteridae, which is the only extant family in the order Phoenicopteriformes. There are four flamingo species distributed throughout the Americas ...
s do today. However, in 2006 an analysis by
Lü Junchang Lü Junchang (; 1965 – 9 October 2018) was a Chinese palaeontologist and professor at the Institute of Geology, Chinese Academy of Geological Sciences. An expert on Mesozoic reptiles, he described and named dozens of dinosaur and pterosaur taxa ...
had as outcome that ''Feilongus'' was not an archaeopterodactyloid, but a member of the
Ornithocheiroidea Ornithocheiroidea (or ornithocheiroids) is a group of pterosaurs within the extinct suborder Pterodactyloidea. They were typically large pterosaurs that lived from the Early to Late Cretaceous periods (Valanginian to Maastrichtian stages), with f ...
''sensu'' Kellner, closer to the
Anhangueridae Anhangueridae is a group of pterosaurs within the suborder Pterodactyloidea. They were among the last pterosaurs to possess teeth. A recent study discussing the group considered the Anhangueridae to be typified by a premaxillary crest and a later ...
. This means that using the alternative terminology of
David Unwin David (; , "beloved one") (traditional spelling), , ''Dāwūd''; grc-koi, Δαυΐδ, Dauíd; la, Davidus, David; gez , ዳዊት, ''Dawit''; xcl, Դաւիթ, ''Dawitʿ''; cu, Давíдъ, ''Davidŭ''; possibly meaning "beloved one". w ...
they are close to the Ornithocheiroidea ''sensu'' Unwin, a group the members of which are typically more adapted to
soaring Soaring may refer to: * Gliding, in which pilots fly unpowered aircraft known as gliders or sailplanes * Lift (soaring), a meteorological phenomenon used as an energy source by some aircraft and birds * ''Soaring'' (magazine), a magazine produced ...
and a
piscivore A piscivore () is a carnivorous animal that eats primarily fish. The name ''piscivore'' is derived . Piscivore is equivalent to the Greek-derived word ichthyophage, both of which mean "fish eater". Fish were the diet of early tetrapod evoluti ...
, or fish-eating, diet. Another publication following this general line of thought has put ''Feilongus'' and ''
Boreopterus ''Boreopterus'' is a genus of boreopterid pterodactyloid pterosaur from the Barremian-Aptian-age Lower Cretaceous Yixian Formation of Dalian, Liaoning, China. Etymology ''Boreopterus'' was named in 2005 by Lü Junchang and Ji Qiang. The type spe ...
'' into a new ornithocheiroid family, the
Boreopteridae Boreopteridae (meaning "northern wings") is a group of pterodactyloid pterosaurs from the Aptian-age Lower Cretaceous Yixian Formation of Liaoning, China. Classification In 2006, Lü Junchang and colleagues named the clade Boreopteridae for the ...
. A 2018 phylogenetic analysis recovers ''Feilongus'' as a
ctenochasmatid Ctenochasmatidae is a group of pterosaurs within the suborder Pterodactyloidea. They are characterized by their distinctive teeth, which are thought to have been used for filter-feeding. Ctenochasmatids lived from the Late Jurassic to the Early C ...
. Specifically, ''Feilongus'' is placed with its
sister taxon In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and t ...
''
Moganopterus ''Moganopterus'' is an extinct genus of ctenochasmatid pterosaur from the Early Cretaceous of western Liaoning Province, China. Discovery and naming The fossil of ''Moganopterus'' was discovered at the village of Xiaosanjiazi near the town of L ...
'' in the subfamily
Moganopterinae Ctenochasmatidae is a group of pterosaurs within the suborder Pterodactyloidea. They are characterized by their distinctive teeth, which are thought to have been used for filter-feeding. Ctenochasmatids lived from the Late Jurassic to the Early C ...
:


See also

*
List of pterosaur genera This list of pterosaurs is a comprehensive listing of all genera that have ever been included in the order Pterosauria, excluding purely vernacular terms. The list includes all commonly accepted genera, but also genera that are now considered inval ...
*
Timeline of pterosaur research This timeline of pterosaur research is a chronologically ordered list of important fossil discoveries, controversies of interpretation, and taxonomic revisions of pterosaurs, the famed flying reptiles of the Mesozoic era. Although pterosaurs w ...


References


External links


''Feilongus''
in The Pterosauria
in The Grave Yard--> {{Portal bar, Paleontology, Cretaceous, China Ctenochasmatoids Early Cretaceous pterosaurs of Asia Taxa named by Alexander Kellner Fossil taxa described in 2005 Yixian fauna Taxa named by Zhou Zhonghe