Fehria

''Bela'' is a genus of sea snails; marine gastropod mollusks in the family Mangeliidae.

Taxonomy

Because of taxonomic uncertainty regarding the type species, many authors in the 19th century e.g. G.O. Sars (1878) and W.H.Dall (1919) used the generic name ''Bela'' for unrelated species correctly placed in ''Propebela'', ''Oenopota'' or ''Curtitoma'', and then used ''Raphitoma'' for species currently placed in ''Bela''.

Species

Species within the genus ''Bela (gastropod)'' include:

* '' Bela alma'' Thiele, 1925
* '' Bela ampla'' Smith, E.A., 1884
* † ''Bela ankae'' Gürs, 2001
* '' Bela anna'' Thiele, 1925
* † ''Bela annemariae'' Lozouet, 2015
* † '' Bela antwerpiensis'' R. Marquet, 1997
* ''Bela atlantidea'' (Knudsen, 1952)
* '' Bela barbiton'' (Melvill, J.C., 1904)
* ''Bela beatriceae'' (Mariottini, 2007)
* † '' Bela belgica'' (Van Regteren Altena, 1959)
* ''Bela bella'' Barnard, 1958
* ''Bela chuni'' Thiele, 1925
* † ''Bela consimilis'' (Harmer, 1915)
* ''Bela cycladensis'' (Reeve, 1845)
* ''Bela decussata'' (Locard, 1897)
* † ''Bela detexta'' (Bellardi, 1877)
* ''Bela dyscrita'' (Watson, R.B., 1881)
* ''Bela erosa'' (Schrenck, 1861)
* † ''Bela fiorentina'' Della Bella, Naldi & Scarponi, 2015
* † ''Bela formica'' Nordsieck, 1977
* ''Bela fuscata'' (Deshayes, 1835)
* † ''Bela gervillei'' (G.P. Deshayes, 1835)
* † ''Bela hispida'' (Bellardi, 1877)
* † ''Bela hispidula'' (Bellardi, 1847)
* † ''Bela keepingi'' (Etheridge & Bell, 1898)
* † ''Bela koeneni'' (Cossmann & Lambert, 1884)
* † ''Bela lirifera'' (Bellardi, 1877)
* '' Bela manolae'' Horro, Gori & Rolán, 2017
* † '' Bela megastoma'' (Brugnone, 1862)
* ''Bela menkhorsti'' van Aartsen, 1988
* ''Bela nassoides'' J. Gardner, 1938
* ''Bela nebula'' ( Montagu, 1803)
* † '' Bela nevropleura'' (Brugnone, 1862)
* † '' Bela nitida'' Pavia, 1976
* ''Bela nuperrima'' (Tiberi, 1855)
* ''Bela obliquigradata'' E.A. Smith, 1884
* ''Bela oceanica'' (Locard, 1897)
* † '' Bela odhneri'' Harmer, 1915
* '' Bela orientalis'' K.H.J. Thiele, 1925
* '' Bela patagonica'' (d'Orbigny, 1841)
* † '' Bela plagisculpta'' Della Bella, Naldi & Scarponi, 2015
* † '' Bela plicatella'' (Bellardi, 1847)
* ''Bela plicatilis'' (Risso, 1826)
* ''Bela powisiana'' (Dautzenberg, 1887)
* † '' Bela proxima'' (Cocconi, 1873)
* † ''Bela pseudoappeliusi'' Naldi, Della Bella & Scarponi, 2013
* † '' Bela pseudoexilis'' Della Bella, Naldi & Scarponi, 2015
* † '' Bela scalariformis'' (Brugnone, 1862)
* '' Bela schoenherri'' Horro, Ryall & Rolán, 2018
* † ''Bela seyithasanensis'' B.M. Landau & al., 2013
* '' Bela simplicata'' É.A.A. Locard, 1896
* ''Bela submarginata'' (G. Bonelli in L.M.D. Bellardi, 1847)
* ''Bela taprurensis'' (Pallary, 1904)
* † ''Bela tenuistriata'' (A. Bell, 1871)
* † '' Bela trinacria'' Mariottini & Smriglio, 2009
* ''Bela turcica'' K.H.J. Thiele, 1925
* ''Bela turgida'' (L.A. Reeve, 1844)
* † ''Bela varovtsiana'' Scarponi & al., 2016
* † ''Bela vulpecula'' (Brocchi, 1814)
* '' Bela zenetouae'' (van Aartsen, 1988)
* ''Bela zonata'' (Locard, 1892)

;Species brought into synonymy :
* ''Bela abyssicola'' (Reeve, 1844): synonym of '' Pleurotoma abyssicola'' Reeve, 1844
* ''Bela abyssorum'' Locard, 1897: synonym of '' Gymnobela abyssorum'' (Locard, 1897)
*'' Bela aegeensis'' (Reeve, 1844): synonym of ''Bela nebula'' ( Montagu, 1803)
* ''Bela africana'' Ardovini, 2004: synonym of ''Sorgenfreispira africana'' (Ardovini, 2004)
* ''Bela africana'' Ardovini, 2008: synonym of ''Sorgenfreispira africana'' (Ardovini, 2004)
* ''Bela alberti'' (Dautzenberg & Fisher, 1906): synonym of ''Phymorhynchus alberti'' (Dautzenberg & Fischer, 1906)
* ''Bela albrechti'' Krause, 1886: synonym of '' Granotoma albrechti'' (Krause, 1886)
* ''Bela aleutica'' (W.H. Dall, 1871): synonym of ''Oenopota aleutica'' (Møller, 1842)
* ''Bela anderssoni'' Strebel, 1908: synonym of ''Falsimohnia anderssoni'' (Strebel, 1908)
* ''Bela angulosa'' Sars G. O., 1878: synonym of '' Propebela angulosa'' (G. O. Sars, 1878)
* ''Bela antarctica'' Strebel, 1908: synonym of ''Conorbela antarctica'' (Strebel, 1908)
* ''Bela apollinea'' (Melvill, J.C., 1904): synonym of ''Leiocithara apollinea'' (Melvill, 1904)
* ''Bela arctica'' A. Adams, 1855: synonym of '' Propebela arctica'' (A. Adams, 1855)
* ''Bela ardovinii'' Mariottini & Oliverio, 2008: synonym of ''Sorgenfreispira ardovinii'' (Mariottini & Oliverio, 2008)
* ''Bela assimilis'' Sars G. O., 1878: synonym of '' Propebela assimilis'' (Sars G. O., 1878)
* ''Bela australis'' Adams & Angas, 1864: synonym of ''Guraleus australis'' (Adams & Angas, 1864)
* ''Bela bergensis'' Friele, 1886: synonym of '' Propebela bergensis'' (Friele, 1886)
* ''Bela bicarinata'' (Couthouy, 1838): synonym of '' Curtitoma violacea'' (Mighels & C. B. Adams, 1842)
* ''Bela blakei'' A. E. Verrill, 1885: synonym of ''Mohnia blakei'' (A. E. Verrill, 1885)
* ''Bela blaneyi'' Bush, 1909: synonym of ''Oenopota blaneyi'' (Bush, 1909)
* ''Bela brachystoma'' (Philippi, 1844): synonym of ''Sorgenfreispira brachystoma'' (Philippi, 1844)
* ''Bela cancellata'' Mighels & Adams, 1842 sensu G. O. Sars, 1878: synonym of ''Bela sarsii'' Verrill, 1880: synonym of ''Oenopota impressa'' (Mörch, 1869)
* ''Bela cancellata'' (Mighels & Adams, 1842): synonym of ''Propebela cancellata'' (Mighels & Adams, 1842)
* † ''Bela candida'' Yokoyama, 1926: synonym of '' Oenopota candida'' (Yokoyama, 1926)
* ''Bela clarae'' Peñas & Rolán, 2008: synonym of ''Bela atlantidea'' (Knudsen, 1952)
* ''Bela concinnula'' Verrill, 1882: synonym of '' Propebela concinnula'' (A. E. Verrill, 1882)
* ''Bela confusa'' (Locard, 1897): synonym of ''Bela brachystoma'' (Philippi, 1844)
* ''Bela conoidea'' Sars G. O., 1878: synonym of ''Curtitoma conoidea'' (Sars G. O., 1878)
* ''Bela costulata'' ( Risso, 1826): synonym of ''Mangelia costulata'' Risso, 1826
* ''Bela decussata'' (Couthouy, 1839): synonym of ''Curtitoma decussata'' (Couthouy, 1839)
* ''Bela demersa'' Tiberi, 1868: synonym of ''Taranis moerchii'' (Malm, 1861)
* ''Bela detegata'' Locard, 1897: synonym of '' Propebela bergensis'' (Friele, 1886)
* ''Bela erythraea'' Jousseaume, 1895: synonym of ''Taranidaphne hongkongensis'' (Sowerby III, 1889)
* ''Bela eva'' Thiele, 1925: synonym of ''Maoritomella eva'' (Thiele, 1925)
* ''Bela exilis'' Ardovini, 2004: synonym of ''Sorgenfreispira exilis'' (Ardovini, 2004)
* ''Bela expansa'' Sars G. O., 1878: synonym of ''Lusitanops expansa'' (Sars G. O., 1878): synonym of ''Lusitanops expansus'' (Sars G. O., 1878)
* ''Bela exquisita'' Yokoyama, 1926: synonym of ''Curtitoma exquisita'' (Yokoyama, 1926)
* ''Bela eva'' Thiele, 1925: synonym of ''Maoritomella eva'' (Thiele, 1925)
* ''Bela excurvata'' Carpenter, 1864: synonym of ''Oenopota excurvata'' (Carpenter, 1864)
* ''Bela expansa'' Sars G. O., 1878: synonym of '' Lusitanops expansa'' (Sars G. O., 1878)
* ''Bela fidicula'' (Gould, 1849): synonym of '' Propebela fidicula'' (Gould, 1849)
* ''Bela filicinctus'' (E. A. Smith, 1882): synonym of ''Horaiclavus filicinctus'' (E. A. Smith, 1882)
* ''Bela fulvicans'' Strebel, 1908: synonym of ''Falsimohnia fulvicans'' (Strebel, 1908)
* ''Bela furfuraculata'' Locard, 1897: synonym of '' Propebela bergensis'' (Friele, 1886)
* ''Bela gazellae'' Strebel, 1905: synonym of ''Mangelia gazellae'' (Strebel, 1905)
* ''Bela ginnania'' (Risso, 1826): synonym of ''Haedropleura septangularis'' (Montagu, 1803)
* ''Bela ginnania'' ( Risso, 1826):B. Otztürk, ''Comments on Shells of Mollusca collected from the Seas of Turkey'', Turk J Zool, 29, pp. 111-112, 2005
synonym of ''Haedropleura septangularis'' (Montagu, 1803)
* ''Bela glacialis'' Thiele, 1912: synonym of '' Lorabela glacialis'' (Thiele, 1912)
* ''Bela gouldii'' Verrill, 1882: synonym of ''Propebela rugulata'' (Reeve, 1846)
* ''Bela graphica'' Locard, 1897: synonym of ''Oenopota graphica'' (Locard, 1897)
* ''Bela grimaldii'' Dautzenberg, 1889: synonym of ''Amphissa acutecostata'' (Philippi, 1844)
* ''Bela grippi'' Dall, 1908: synonym of ''Bellaspira grippi'' (Dall, 1908)
* ''Bela guernei'' Dautzenberg, 1891: synonym of ''Belomitra quadruplex'' (Watson, 1882)
* ''Bela harpa'' Dall, 1885: synonym of ''Oenopota harpa'' (Dall, 1885)
* ''Bela hebes'' Verrill, 1880: synonym of ''Curtitoma hebes'' (Verrill, 1880)
* ''Bela holomera'' Locard, 1897: synonym of ''Gymnobela pyrrhogramma'' (Dautzenberg & Fischer, 1896)
* ''Bela iessoensis'' Smith E. A., 1875: synonym of '' Obesotoma iessoensis'' (Smith E. A., 1875)
* ''Bela incisula'' Verrill, 1882: synonym of ''Curtitoma incisula'' (Verrill, 1882)
* ''Bela kobelti'' Verkrüzen, 1876: synonym of '' Granotoma kobelti'' (Verkrüzen, 1876)
* ''Bela koreni'' Friele, 1886: synonym of ''Oenopota koreni'' (Friele, 1886)
* ''Bela krausei'' Dall, 1887: synonym of '' Granotoma krausei'' (Dall, 1887)
* ''Bela laevigata'' (Philippi, 1836): synonym of ''Bela zonata'' (Locard, 1892)
* ''Bela laevigata'' Dall, 1871: synonym of '' Obesotoma laevigata'' (Dall, 1871)
* ''Bela limatula'' Locard, 1896: synonym of ''Amphissa acutecostata'' (Philippi, 1844)
* ''Bela luetkeana'' Krause, 1885: synonym of ''Propebela luetkeana'' (Krause, 1885)
* ''Bela lütkeana'' Krause, 1885: synonym of ''Bela luetkeana'' Krause, 1885: synonym of ''Propebela luetkeana'' (Krause, 1885)
* ''Bela magellanica'' (Martens, 1881): synonym of ''Oenopota magellanica'' (Martens, 1881)
* ''Bela martensi'' Strebel, 1905: synonym of ''Mangelia martensi'' (Strebel, 1905)
* ''Bela metschigmensis'' Krause, 1886: synonym of '' Oenopota metschigmensis'' (Krause, 1886)
* ''Bela michaelseni'' Strebel, 1905: synonym of ''Mangelia michaelseni'' (Strebel, 1905)
* ''Bela mingoranceae'' Martin Perez & Vera-Pelaez, 2006: synonym of ''Bela powisiana'' (Dautzenberg, 1887)
* ''Bela minuscularia'' Locard, 1897: synonym of '' Curtitoma violacea'' (Mighels & C. B. Adams, 1842)
* ''Bela mitralis'' Adams & Angas, 1864: synonym of ''Guraleus mitralis'' (Adams & Angas, 1864)
* ''Bela mitrula'' Lovén, 1846: synonym of '' Propebela exarata'' (Møller, 1842)
* ''Bela murdochiana'' Dall, 1885: synonym of ''Oenopota murdochiana'' (Dall, 1885)
* ''Bela neozelanica'' Suter, 1908: synonym of '' Scrinium neozelanica'' (Suter, 1908)
* ''Bela nobilis'' (Møller, 1842): synonym of ''Propebela nobilis'' (Møller, 1842)
* ''Bela notophila'' Strebel, 1908: synonym of '' Lorabela notophila'' (Strebel, 1908)
* ''Bela obliqua'' Sars G. O., 1878: synonym of '' Oenopota obliqua'' (Sars G.O., 1878)
* ''Bela optima'' Thiele, 1925: synonym of ''Microdrillia optima'' (Thiele, 1925)
* ''Bela ornata'' (Locard, 1891): synonym of ''Mangelia costulata'' Risso, 1826
* † ''Bela paessleri'' H. Strebel, 1905: synonym of ''Mangelia paessleri'' (H. Strebel, 1905)
* ''Bela pelseneri'' Strebel, 1908: synonym of '' Lorabela pelseneri'' (Strebel, 1908)
* ''Bela plicatula'' Thiele, 1912: synonym of ''Lorabela plicatula'' (Thiele, 1912)
* ''Bela polysarca'' (Dautzenberg & Fischer H., 1896): synonym of ''Gymnobela frielei'' (Verrill, 1885)
* ''Bela purissima'' Strebel, 1908: synonym of '' Typhlodaphne purissima'' (Strebel, 1908)
* ''Bela pygmaea'' Verrill, 1882: synonym of ''Curtitoma ovalis'' (Friele, 1877)
* ''Bela pyramidalis'' (Ström, 1788): synonym of ''Oenopota pyramidalis'' (Strøm, 1788)
* ''Bela rathbuni'' Verrill, 1882: synonym of ''Propebela rathbuni'' (Verrill, 1882)
* ''Bela recondita'' Locard, 1897: synonym of ''Gymnobela pyrrhogramma'' (Dautzenberg & Fischer, 1896)
* ''Bela regina'' Thiele, 1925: synonym of '' Tomopleura regina'' (Thiele, 1925)
* ''Bela robusta'' Packard, 1866: synonym of ''Obesotoma robusta'' (Packard, 1866)
* ''Bela rugulata'' (Reeve, 1846): synonym of ''Propebela rugulata'' (Reeve, 1846)
* ''Bela sansibarica'' Thiele, 1925: synonym of ''Microdrillia sansibarica'' (Thiele, 1925)
* ''Bela sarsii'' Verrill, 1880: synonym of ''Oenopota impressa'' (Mörch, 1869)
* ''Bela scalaris'' (Møller, 1842): synonym of ''Propebela scalaris'' (Møller, 1842)
* ''Bela scalaroides'' Sars G. O., 1878: synonym of ''Propebela scalaroides'' (Sars G. O., 1878)
* ''Bela schmidti'' Friele, 1886: synonym of ''Oenopota harpa'' (Dall, 1885)
* ''Bela sculpturata'' Dall, 1887: synonym of ''Mangelia sculpturata'' (Dall, 1887)
* ''Bela septangularis'' (Montagu, 1803): synonym of ''Haedropleura septangularis'' (Montagu, 1803)
* ''Bela septenvillei'' (Dautzenberg & Durouchoux, 1913): synonym of ''Bela nebula'' ( Montagu, 1803)
* ''Bela solida'' Dall, 1887: synonym of '' Obesotoma solida'' (Dall, 1887)
* ''Bela spitzbergensis'' Friele, 1886: synonym of ''Propebela spitzbergensis'' (Friele, 1886)
* ''Bela striata'' Hutton, 1873: synonym of †''Iredalula striata'' (Hutton, 1873)
* ''Bela striatula'' Thiele, 1912: synonym of ''Belalora striatula'' (Thiele, 1912)
* ''Bela subarctica'' Derjugin, 1924: synonym of ''Propebela rugulata'' (Reeve, 1846)
* ''Bela subturgida'' Verrill, 1884: synonym of '' Propebela subturgida'' (Verrill, 1884)
* ''Bela subvitrea'' Verrill, 1884: synonym of ''Propebela subvitrea'' (Verrill, 1884)
* ''Bela tenuicostata'' Sars G. O., 1878: synonym of ''Oenopota tenuicostata'' (Sars G.O., 1878)
* ''Bela tenuilirata'' Dall, 1871: synonym of '' Obesotoma tenuilirata'' (Dall, 1871)
* ''Bela tenuilirata'' Krause, 1886: synonym of ''Curtitoma lawrenciana'' (Dall, 1919)
* ''Bela tumida'' Posselt, 1898: synonym of '' Obesotoma tumida'' (Posselt, 1898)
* ''Bela turricula'' (Montagu, 1803): synonym of '' Propebela turricula'' (Montagu, 1803)
* ''Bela turriculata'' Locard, 1892: synonym of '' Propebela turricula'' (Montagu, 1803)
* ''Bela turrita'' Strebel, 1908: synonym of ''Belaturricula turrita'' (Strebel, 1908)
* ''Bela violacea'' (Mighels & Adams, 1842): synonym of '' Curtitoma violacea'' (Mighels & C. B. Adams, 1842)
* ''Bela woodiana'' (Møller, 1842): synonym of '' Obesotoma woodiana'' (Møller, 1842)
* ''Bela yanamii'' Yokoyama, 1926: synonym of ''Plicifusus yanamii'' (Yokoyama, 1926)
* ''Bela zonatum'' (Locard, 1891): duplicate of ''Bela zonata''

; Nomina dubia:
* '' Bela fortis'' (Reeve, 1844): ''nomen dubium''
* '' Bela minuta'' (Reeve, 1844): ''nomen dubium''
* ''Bela turgida'' (Reeve, 1844): ''nomen dubium''

References

* Vaught, K.C. (1989). ''A classification of the living Mollusca''. American Malacologists: Melbourne, FL (USA). . XII, 195 pp
* Howson, C.M.; Picton, B.E. (Ed.) (1997). ''The species directory of the marine fauna and flora of the British Isles and surrounding seas''. Ulster Museum Publication, 276. The Ulster Museum: Belfast, UK. . vi, 508
* Gofas, S.; Le Renard, J.; Bouchet, P. (2001). ''Mollusca, in: Costello, M.J. et al. (Ed.) (2001). European register of marine species: a check-list of the marine species in Europe and a bibliography of guides to their identification''. Collection Patrimoines Naturels, 50: pp. 180–213
* W. Baluk. 2003. ''Middle Miocene (Badenian) gastropods from Korytnica, Poland; Part IV – Turridae''. Acta Geological Polonica 53(1):29-78
* Mariottini P., Smriglio C., Di Giulio A. & Oliverio M. 2009. A'' new fossil conoidean from the Pliocene of Italy, with comments on the Bela menkhorsti complex (Gastropoda: Conidae).'' Journal of Conchology 40(1): 5-14

External links

Gray J.E. (1847). ''On the classification of the British Mollusca by W E Leach''. Annals and Magazine of Natural History. ser. 1, 20: 267-273

P Bartsch. "The Nomenclatorial Status of Certain Northern Turritid Mollusks"; Proceedings of the biological Society of Washington 54, 1-14, 1941

Della Bella G., Naldi F. & Scarponi D. (2015). ''Molluschi marini del Plio-Pleistocene dell'Emilia-Romagna e della Toscana - Superfamiglia Conoidea, vol. 4, Mangeliidae II''. Lavori della Società Italiana di Malacologia. 26: 1-80



Tucker, J.K. 2004 ''Catalog of recent and fossil turrids (Mollusca: Gastropoda)''. Zootaxa 682:1-1295

{{DEFAULTSORT:Bela (Gastropod)

Gastropod genera
Taxa named by John Edward Gray