Faxinalipterus
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''Faxinalipterus'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of ornithodiran
archosaur Archosauria () is a clade of diapsids, with birds and crocodilians as the only living representatives. Archosaurs are broadly classified as reptiles, in the cladistic sense of the term which includes birds. Extinct archosaurs include non-avian d ...
, originally described as a
pterosaur Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to ...
, from the
Late Triassic The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch ...
Caturrita Formation of southern
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
. A study from 2022 reinterpreted the fossil remains and instead suggests them to belong to a member of the Lagerpetidae, a more basal clade of pterosauromorph.


Description

The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
, ''Faxinalipterus minima'' was named and described in 2010 by
José Fernando Bonaparte José is a predominantly Spanish and Portuguese form of the given name Joseph. While spelled alike, this name is pronounced differently in each language: Spanish ; Portuguese (or ). In French, the name ''José'', pronounced , is an old vernacul ...
,
Marina Bento Soares A marina (from Spanish , Portuguese and Italian : ''marina'', "coast" or "shore") is a dock or basin with moorings and supplies for yachts and small boats. A marina differs from a port in that a marina does not handle large passenger ship ...
and César Leandro Schultz. The generic name is derived from
Faxinal do Soturno Faxinal do Soturno is a municipality in the state of Rio Grande do Sul, Brazil. See also *List of municipalities in Rio Grande do Sul This is a list of the municipalities in the state of Rio Grande do Sul (RS), located in the South Region of Br ...
and a Latinised Greek ''pteron'', "wing". The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
, from
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
''minima'', means "smallest". The name was later amended and changed to ''minimus'' in accordance with the rules of the International Commission on Zoological Nomenclature (ICZN). Fossils of the species were found in 2002 and 2005 at a site 1.5 kilometres northeast of Faxinal do Soturno,
Rio Grande do Sul Rio Grande do Sul (, , ; "Great River of the South") is a Federative units of Brazil, state in the South Region, Brazil, southern region of Brazil. It is the Federative_units_of_Brazil#List, fifth-most-populous state and the List of Brazilian st ...
, from the Caturrita Formation dating from the Carnian-
Norian The Norian is a division of the Triassic Period. It has the rank of an age (geochronology) or stage (chronostratigraphy). It lasted from ~227 to million years ago. It was preceded by the Carnian and succeeded by the Rhaetian. Stratigraphic defi ...
, 220–215 million years old. The
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
, UFRGS PV0927T, is part of the collection of the Universidade Federal do Rio Grande do Sul and consists of several, partially fragmentary, limb elements, perhaps of a single individual. These do not include bones from the hand. An upper jaw fragment, a left
maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The t ...
with three teeth, has been referred to the species, but later determined to represent an animal distinct from ''Faxinalipterus'', which was named ''
Maehary ''Maehary'' (meaning "one who looks to the sky" in Guaraní) is a genus of pterosauromorph from the Late Triassic (Norian) Caturrita Formation of Rio Grande do Sul, Brazil. The genus contains a single species, ''M. bonapartei'', known from a par ...
''. ''Faxinalipterus'' is a rather small animal, with an estimated total
humerus The humerus (; ) is a long bone in the arm that runs from the shoulder to the elbow. It connects the scapula and the two bones of the lower arm, the radius and ulna, and consists of three sections. The humeral upper extremity consists of a roun ...
length of eighteen millimetres. The size has been compared to that of a
sparrow Sparrow may refer to: Birds * Old World sparrows, family Passeridae * New World sparrows, family Passerellidae * two species in the Passerine family Estrildidae: ** Java sparrow ** Timor sparrow * Hedge sparrow, also known as the dunnock or hedg ...
. The hind limbs, much better conserved than the front limbs, are relatively long.


Taxonomy

The describers assigned ''Faxinalipterus'' to the Pterosauria, based on its long hollow limbs and saddle-shaped upper joint of the relatively short and robust humerus, suitable to perform a wing stroke. They saw it as perhaps the oldest pterosaur known, as it possibly predates European finds from the Norian. they see that the possible age difference between pterosaurs cannot be large as an indication of rapid evolution in early pterosaurs. Because the Caturrita Formation consists of terrestrial sandstones, that evolution would have had its origins in a terrestrial, not coastal, habitat. They also concluded ''Faxinalipterus'' is the most basal known pterosaur, basal features including a lack of fusion between
tibia The tibia (; ), also known as the shinbone or shankbone, is the larger, stronger, and anterior (frontal) of the two bones in the leg below the knee in vertebrates (the other being the fibula, behind and to the outside of the tibia); it connects ...
and
fibula The fibula or calf bone is a leg bone on the lateral side of the tibia, to which it is connected above and below. It is the smaller of the two bones and, in proportion to its length, the most slender of all the long bones. Its upper extremity is ...
, a thin
radius In classical geometry, a radius ( : radii) of a circle or sphere is any of the line segments from its center to its perimeter, and in more modern usage, it is also their length. The name comes from the latin ''radius'', meaning ray but also the ...
and a coracoid that has not fused to the
scapula The scapula (plural scapulae or scapulas), also known as the shoulder blade, is the bone that connects the humerus (upper arm bone) with the clavicle (collar bone). Like their connected bones, the scapulae are paired, with each scapula on eithe ...
. However, Alexander Kellner has suggested ''Faxinalipterus'' might be not be a pterosaur but a basal member of the
Pterosauromorpha Pterosauromorpha (meaning "pterosaur-like forms") is one of the two basic divisions of Ornithodira that includes pterosaurs and all taxa that are closer to them than to dinosaurs and their close relatives (i.e. Dinosauromorpha). In addition to pt ...
instead or, if the lack of fusion between tibia and fibula is
plesiomorphic In phylogenetics, a plesiomorphy ("near form") and symplesiomorphy are synonyms for an ancestral character shared by all members of a clade, which does not distinguish the clade from other clades. Plesiomorphy, symplesiomorphy, apomorphy, and ...
, even a sister taxon of the Ornithodira. Fabio Marco Dalla Vecchia (2013) stated that he was "unable to find any unequivocal pterosaur features" in the known fossils of ''F. minima''; in fact, according to the author, "the purported humerus is quite unlike the humeri of the Triassic pterosaurs". Dalla Vecchia did not consider ''Faxinalipterus'' to be a pterosaur, but did not state what group of vertebrates it belonged to. ''Faxinalipterus'' was reinterpreted as an early member of Lagerpetidae by Kellner and colleagues in 2022, who also removed the maxilla from the referred specimens and assigned it to the newly named basal pterosauromorph ''
Maehary ''Maehary'' (meaning "one who looks to the sky" in Guaraní) is a genus of pterosauromorph from the Late Triassic (Norian) Caturrita Formation of Rio Grande do Sul, Brazil. The genus contains a single species, ''M. bonapartei'', known from a par ...
''.


See also

*
2010 in paleontology Plants Angiosperms Molluscs Newly named bivalves Arthropods Fishes Amphibians Newly named amphibians Basal reptiles Newly named basal reptiles Ichthyopterygians Newly named ichthyopterygians Lepidosauromorphs Newly name ...


References


External links


Pterosaur Database entry for ''Faxinalipterus''
{{Taxonbar, from=Q735037 Late Triassic reptiles Late Triassic reptiles of South America Triassic Brazil Fossils of Brazil Paraná Basin Fossil taxa described in 2010 Taxa named by José Bonaparte Lagerpetidae