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''Macroprotodon cucullatus'', commonly known as the false smooth snake, is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of mildly
venom Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a st ...
ous
colubrid Colubridae (, commonly known as colubrids , from la, coluber, 'snake') is a family of snakes. With 249 genera, it is the largest snake family. The earliest species of the family date back to the Oligocene epoch. Colubrid snakes are found on ev ...
snake Snakes are elongated, limbless, carnivorous reptiles of the suborder Serpentes . Like all other squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales. Many species of snakes have skulls with several more j ...
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...
to the Mediterranean Basin.


Geographic distribution

''M. cucullatus'' is found in
Algeria ) , image_map = Algeria (centered orthographic projection).svg , map_caption = , image_map2 = , capital = Algiers , coordinates = , largest_city = capital , relig ...
,
Egypt Egypt ( ar, مصر , ), officially the Arab Republic of Egypt, is a transcontinental country spanning the northeast corner of Africa and southwest corner of Asia via a land bridge formed by the Sinai Peninsula. It is bordered by the Medit ...
,
Israel Israel (; he, יִשְׂרָאֵל, ; ar, إِسْرَائِيل, ), officially the State of Israel ( he, מְדִינַת יִשְׂרָאֵל, label=none, translit=Medīnat Yīsrāʾēl; ), is a country in Western Asia. It is situated ...
,
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical ...
,
Libya Libya (; ar, ليبيا, Lībiyā), officially the State of Libya ( ar, دولة ليبيا, Dawlat Lībiyā), is a country in the Maghreb region in North Africa. It is bordered by the Mediterranean Sea to the north, Egypt to the east, Suda ...
,
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria t ...
,
Palestinian Territories The Palestinian territories are the two regions of the former British Mandate for Palestine that have been militarily occupied by Israel since the Six-Day War of 1967, namely: the West Bank (including East Jerusalem) and the Gaza Strip. The ...
,
Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of ...
,
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
, and
Tunisia ) , image_map = Tunisia location (orthographic projection).svg , map_caption = Location of Tunisia in northern Africa , image_map2 = , capital = Tunis , largest_city = capital , ...
.


Description

''Macroprotodon cucullatus'' is a small snake, usually not exceeding in total length (including tail). As the common name implies, the
dorsal scales In snakes, the dorsal scales are the longitudinal series of plates that encircle the body, but do not include the ventral scales In snakes, the ventral scales or gastrosteges are the enlarged and transversely elongated scales that extend down t ...
are smooth, and are arranged in 19-23 rows.
Dorsally Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position prov ...
it is tan or gray, with small brown spots, or with darker and lighter streaks. Ventrally it is yellow to coral-red, uniform or spotted with black, the spots sometimes confluent at the midline. Boulenger, G.A. (1896). ''Catalogue of the Snakes in the British Museum (Natural History), Volume III., Containing the Colubridæ (Opisthoglyphæ and Proteroglyphæ),...'' London: Trustees of the British Museum (Natural History). (Taylor and Francis, printers.) xiv + 727 pp. + Plates I.- XXV. (''Macroprotodon cucullatus'', pp. 175-177, Figure 12). It has a blackish collar which may extend to the top of the head. There is usually a dark streak from the nostril to the corner of the mouth, passing under or through the eye. The eye is rather small, and the pupil, when seen contracted in bright light, is oval. The snout is flattened, and the rostral is wide and low, barely visible from above. The 6th upper labial extends upward and contacts the parietal. There is usually one anterior temporal. Arnold, E.N., and J.A. Burton (1978). ''A Field Guide to the Reptiles and Amphibians of Britain and Europe''. London: Collins. 272 pp. . (''Macroprotodon cucullatus'', pp. 206-207 + Plate 38 + Map 119).


Habitat

The natural
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
s of ''M. cucullatus'' are temperate
forest A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' ...
s, temperate shrubland, subtropical or tropical dry shrubland, Mediterranean-type shrubby vegetation, freshwater
spring Spring(s) may refer to: Common uses * Spring (season) Spring, also known as springtime, is one of the four temperate seasons, succeeding winter and preceding summer. There are various technical definitions of spring, but local usage of ...
, rocky areas, sandy shores,
arable land Arable land (from the la, arabilis, "able to be ploughed") is any land capable of being ploughed and used to grow crops.''Oxford English Dictionary'', "arable, ''adj''. and ''n.''" Oxford University Press (Oxford), 2013. Alternatively, for th ...
, pastureland,
plantation A plantation is an agricultural estate, generally centered on a plantation house, meant for farming that specializes in cash crops, usually mainly planted with a single crop, with perhaps ancillary areas for vegetables for eating and so on. The ...
s, rural gardens, and
urban area An urban area, built-up area or urban agglomeration is a human settlement with a high population density and infrastructure of built environment. Urban areas are created through urbanization and are categorized by urban morphology as cities ...
s.


Behaviour

The false smooth snake is nocturnal in some parts of its range but mainly diurnal in the Balearic Islands.


Diet

''M. cucullatus''
preys Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill th ...
upon small lizards such as geckos and lizards of the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
''
Lacerta Lacerta is one of the 88 modern constellations defined by the International Astronomical Union. Its name is Latin for lizard. A small, faint constellation, it was defined in 1687 by the astronomer Johannes Hevelius. Its brightest stars form a "W" ...
''. Small mammals and nestling birds are also eaten.


Reproduction

Females of ''M. cucullatus'' may breed in alternate years. Two to six eggs are laid in a damp spot under a stone, buried in the soil or hidden in dense vegetation. They hatch in about eight weeks into juveniles with a snout-to-vent length of .


Venom

''M. cucullatus'' possesses a mild venom, which is delivered by means of enlarged grooved teeth in the upper jaw. Although the venom is effective on lizards, this snake is not harmful to humans due to its small size.


Conservation status

The IUCN has listed the false smooth snake as being of "
Least Concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. T ...
". This is because it has a wide distribution and a large population, seems to be tolerant of some habitat modification and its population is unlikely to be declining fast enough to qualify it for listing in a more threatened category.


See also

* List of reptiles of Italy


References


Further reading

*Busack, S.D., and C.J. McCoy (1990). "Distribution, variation, and biology of ''Macroprotodon cucullatus'' (Reptilia: Colubridae: Boiginae)". ''Annals of Carnegie Museum'' 59 (4): 261-286. * Geoffroy Saint-Hilaire, .(1827). "''Description des reptiles qui se trouvent en Égypte'' ". pp. 115–160 + plates. ''In'': Savigny, M.J.C. (1809-1827). ''Description de l'Égypte ...'' olume 24 Paris: C.L.F. Panckoucke. (Second Edition, 1829). (''Coluber cucullatus'', new species, pp. 148, 151 + Plate 8, figures 3 & 3').


External links

*Corti, C.; Mellado, V.P.; Geniez, P.; El Din, S.B.; Martinez-Solano, I; Sindaco, R.; Romano, A. (2009). IUCN (International Union for Conservation of Nature and Natural Resources) Red List Assessment, Version 3.1. http://eol.org/data_objects/18923870. Accessed October 2013.
''Macroprotodon cucullatus'' (Geoffroy Saint-Hilaire, 1827)
. {{Taxonbar, from=Q1728800 Macroprotodon Snakes of Africa Snakes of Asia Reptiles of Europe Fauna of the Middle East Reptiles of North Africa Reptiles described in 1827 Taxa named by Isidore Geoffroy Saint-Hilaire Taxonomy articles created by Polbot