Falco Chicquera
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The red-necked falcon (''Falco chicquera'') is a
bird of prey Birds of prey or predatory birds, also known as raptors, are hypercarnivorous bird species that actively hunt and feed on other vertebrates (mainly mammals, reptiles and other smaller birds). In addition to speed and strength, these predators h ...
in the
falcon Falcons () are birds of prey in the genus ''Falco'', which includes about 40 species. Falcons are widely distributed on all continents of the world except Antarctica, though closely related raptors did occur there in the Eocene. Adult falcons ...
family with two disjunct populations, one in India and the other in Africa. This medium-sized falcon has bluish grey wings and upper body, a chestnut red cap with short chin straps passing through the eye. The primary feathers of the wing are black and a single black band at the tip of the tail are distinctive. The Indian subspecies ''Falco chicquera chicquera'' also known as the red-headed merlin or red-headed falcon is found mainly in the open plains of the India Subcontinent although it is thought to have occurred further west in southeastern Iran. The subspecies ''Falco chicquera ruficollis'' found in sub-Saharan Africa is sometimes treated as a full species, the rufous-necked falcon (''Falco ruficollis''), on the basis of its well-separated geographic range and distinctive pattern. It appears very similar to the Indian form but has dark barring on the upperparts, a rufous breast band, and black moustachial and eye stripes. As in most falcons, the females are larger and falconers in India called the female ''turumti'' and the male as ''chatwa''. They hunt in pairs mostly at dawn and dusk, capturing small birds, bats and squirrels.


Description

The red-necked falcon is a medium-sized, long-winged
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
with a bright rufous crown and nape. It is on average 30–36 cm in length with a wingspan of 85 cm. The wings and upper parts are bluish grey and the tail has narrow bars, a broad subterminal black band tipped with white. The wingtip does not reach the tip of the tail at rest. The second and third primaries are the longest and almost equal in length while the first is a fourth the length. The first two primaries are notched. The legs, ceres and eyering are yellow. The tip of the bill is black while the basal portion is greenish yellow. The voice of this species is a shrill ''ki-ki-ki-ki''. The sexes are similar except in size, males are smaller than females as is usual in falcons. Young birds are buff below with less extensive barring and duller upper plumage.


Subspecies

The adult of the African
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
''Falco chicquera ruficollis'' (a full species, ''F. ruficollis'', in many treatments) first described by William Swainson in 1837 has a white face apart from black on the moustachial stripe. The upperparts are pale grey, with black primary wing feathers and the broad tail band. The underparts are white with dark barring on the underwings, lower breast, belly and undertail. There is a rufous foreneck band (not present in the Indian form). West African males are known to weigh between 139 and 178 grams, while females are found between 190 and 305 grams. The particularly large African birds from south of the
Zambezi River The Zambezi River (also spelled Zambeze and Zambesi) is the fourth-longest river in Africa, the longest east-flowing river in Africa and the largest flowing into the Indian Ocean from Africa. Its drainage basin covers , slightly less than hal ...
are often separated as subspecies ''Falco chicquera horsbrughi'', which was described by
J. W. B. Gunning Jan Willem Boudewijn Gunning (3 September 1860 in Hilversum, North Holland – 26 June 1913 in Pretoria), was a Dutch physician, who served as the director of both the Transvaal Museum, Staatsmuseum and what was then known as the National Zoo ...
&
Austin Roberts Austin Roberts may refer to: * Austin Roberts (American football) (born 1995), American football tight end *Austin Roberts (singer) (born 1945), American singer and songwriter *Austin Roberts (zoologist) Austin Roberts (3 January 1883 – 5 May ...
in 1911 on the basis of a single specimen, but the size variation may be clinal and the subspecies may not be valid. The Asian
nominate subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
''Falco chicquera chicquera'' has rufous moustachial stripes, lacks the buff breast band, and is less extensively barred than the African form.


Taxonomy

The species was described on the basis of a specimen obtained by
François Levaillant François Levaillant (born Vaillant, later in life as Le Vaillant, ''"The Valiant"'') (6 August 1753 – 22 November 1824) was a French author, explorer, naturalist, zoological collector, and noted ornithologist. He described many new species of ...
from Chandernagore in Bengal where he was told it went by the name of Chicquera. This local name was probably attributed to this species by mistake as the name probably referred to a
shikra The shikra (''Accipiter badius'') is a small bird of prey in the family Accipitridae found widely distributed in Asia and Africa where it is also called the little banded goshawk. The African forms may represent a separate species but have usual ...
. Levaillant did not use binomials but called it Le Chicquera and
François Marie Daudin François Marie Daudin (; 29 August 1776 in Paris – 30 November 1803 in Paris) was a French zoologist. With legs paralyzed by childhood disease, he studied physics and natural history, but ended up being devoted to the latter. Daudin wrote ' ( ...
gave it the binomial ''Falco chicquera'' in 1800. Early authors placed it in the genus ''Aesalon'' along with the
merlin Merlin ( cy, Myrddin, kw, Marzhin, br, Merzhin) is a mythical figure prominently featured in the legend of King Arthur and best known as a mage, with several other main roles. His usual depiction, based on an amalgamation of historic and le ...
and later in the genus ''Lithofalco''. Others have placed it in the genus ''Chicquera'' or ''Hypotriorchis''. Studies of molecular sequence divergence in the
cytochrome b Cytochrome b within both molecular and cell biology, is a protein found in the mitochondria of eukaryotic cells. It functions as part of the electron transport chain and is the main subunit of transmembrane cytochrome bc1 and b6f complexes. F ...
gene suggest that the African and Indian forms are distinctive due to separation for as long as 0.9 million years. With non-overlapping ranges they are often treated as full species.


Habitat and distribution

In Africa, the red-necked falcon is found in
semi-desert A semi-arid climate, semi-desert climate, or steppe climate is a dry climate sub-type. It is located on regions that receive precipitation below potential evapotranspiration, but not as low as a desert climate. There are different kinds of semi-ar ...
,
savannah A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the Canopy (forest), canopy does not close. The open canopy allows sufficient light to rea ...
and other dry open country with some trees, but also riverine forest. It often perches and makes use of the crowns of Borassus palms (''
Borassus aethiopum ''Borassus aethiopum'' is a species of '' Borassus'' palm from Africa. In English it is variously referred to as African fan palm, African palmyra palm, deleb palm, ron palm, toddy palm, black rhun palm, rônier palm (from the French). It is wide ...
'') for breeding. They are mostly resident but may make nomadic movements in response to weather. In India, they are found in open habitats and is not found in dense forests or high hills. The nominate subspecies is believed to have occurred west until Iran although no records exist from there since 1970. They were recorded breeding in 1911 by
Nikolai Zarudny Nikolai Alekseyvich Zarudny (russian: Николай Алексеевич Зарудный; rus, Николай Алексеевич Зарудный, r=Nikolay Alekseevich Zarudny. His name has been transliterated a number of other ways; especial ...
. It is a winter vagrant in northern Sri Lanka where it was first recorded by E.L. Layard.


Behaviour

The red-necked falcon usually hunts in pairs, often at dawn and dusk, sometimes utilizing a technique in which one of the pair flies low and flushes up small birds while the other follows higher up and seizes the prey as it flushes from cover. They fly with a fast and dashing flight. It prefers to prey on birds found in open areas and some of the species it has been recorded to hunt are
Eurasian tree sparrow The Eurasian tree sparrow (''Passer montanus'') is a passerine bird in the sparrow family with a rich chestnut crown and nape, and a black patch on each pure white cheek. The sexes are similarly plumaged, and young birds are a duller version ...
(''Passer montanus''),
house sparrow The house sparrow (''Passer domesticus'') is a bird of the sparrow family Passeridae, found in most parts of the world. It is a small bird that has a typical length of and a mass of . Females and young birds are coloured pale brown and grey, a ...
(''Passer domesticus''),
white-browed wagtail The white-browed wagtail or large pied wagtail (''Motacilla maderaspatensis'') is a medium-sized bird and is the largest member of the wagtail family. They are conspicuously patterned with black above and white below, a prominent white brow, sho ...
(''Motacilla madaraspatensis''),
rosy starling The rosy starling (''Pastor roseus'') is a passerine bird in the starling family, Sturnidae, also known as the rose-coloured starling or rose-coloured pastor. The species was recently placed in its own monotypic genus, ''Pastor'', and split fro ...
(''Sturnus roseus''),
chestnut-tailed starling The chestnut-tailed starling (''Sturnia malabarica''), also called grey-headed starling and grey-headed myna is a member of the starling family. It is a resident or partially migratory species found in wooded habitats in India and Southeast Asia. ...
(''Sturnus malabaricus''),
Indian cuckoo The Indian cuckoo (''Cuculus micropterus'') is a member of the cuckoo order of birds, the Cuculiformes, that is found in the Indian subcontinent and Southeast Asia. It ranges from India, Bangladesh, Bhutan, Nepal and Sri Lanka east to Indonesia ...
(''Cuculus micropterus''),
Kentish plover The Kentish plover (''Charadrius alexandrinus'') is a small cosmopolitan shorebird (40-44 g) of the family Charadriidae that breeds on the shores of saline lakes, lagoons, and coasts, populating sand dunes, marshes, semi-arid desert, and tundra.Sz ...
(''Charadrius alexandrinus''),
little ringed plover The little ringed plover (''Charadrius dubius'') is a small plover. The genus name ''Charadrius'' is a Late Latin word for a yellowish bird mentioned in the fourth-century Vulgate. It derives from Ancient Greek ''kharadrios'' a bird found in river ...
(''Charadrius dubius''), ashy-crowned finch-lark (''Eremoptrix griseus''), besides robins, quails, babblers, swifts, bulbuls, pipits, larks (mainly ''Calandrella'', ''Alauda'', ''Galerida'' sp.),
pied cuckoo The Jacobin cuckoo (''Clamator jacobinus''), also pied cuckoo or pied crested cuckoo, is a member of the cuckoo order of birds that is found in Africa and Asia. It is partially migratory and in India, it has been considered a harbinger of the mons ...
(''Clamator jacobinus''),
rock pigeon The rock dove, rock pigeon, or common pigeon ( also ; ''Columba livia'') is a member of the bird family Columbidae (doves and pigeons). In common usage, it is often simply referred to as the "pigeon". The domestic pigeon (''Columba livia domes ...
(''Columba livia''),
collared dove ''Streptopelia'' is a genus of birds in the pigeon and dove family Columbidae. These are mainly slim, small to medium-sized species. The upperparts tend to be pale brown and the underparts are often a shade of pink. Many have a characteristic bla ...
(''Streptopelia decaocto''),
laughing dove The laughing dove (''Spilopelia senegalensis'') is a small pigeon that is a resident breeder in Africa, the Middle East, South Asia, and Western Australia where it has established itself in the wild after being released from Perth Zoo in 1898. Th ...
(''Streptopelia senegelensis''),
brown crake The brown crake (''Zapornia akool''), or brown bush-hen, is a waterbird in the rail and crake family (''Rallidae'') found in South Asia. The specific name (zoology), species name, ''akool'', is of uncertain origin. It may come from Hindu mytholo ...
(''Lanius cristatus''),
tailor bird Tailorbirds are small birds, most belonging to the genus ''Orthotomus''. While they were often placed in the Old World warbler family Sylviidae, recent research suggests they more likely belong in the Cisticolidae and they are treated as such in ...
(''Orthotomus sutorius''),
brown shrike The brown shrike (''Lanius cristatus'') is a bird in the shrike family that is found mainly in Asia. It is closely related to the red-backed shrike (''L. collurio'') and isabelline shrike (''L. isabellinus''). The genus name, ''Lanius'', is deri ...
(''Lanius cristatus''),
white-breasted kingfisher The white-throated kingfisher (''Halcyon smyrnensis'') also known as the white-breasted kingfisher is a tree kingfisher, widely distributed in Asia from the Sinai east through the Indian subcontinent to China and Indonesia. This kingfisher is a ...
(''Halcyon smyrnensis''),
little stint The little stint (''Calidris minuta'' or ''Erolia minuta''), is a very small wader. It breeds in arctic Europe and Asia, and is a long-distance migrant, wintering south to Africa and south Asia. It occasionally is a vagrant to North America a ...
(''Calidris minuta''), plain martin (''Riparia paludicola'') and
pied bushchat The pied bush chat (''Saxicola caprata'') is a small passerine bird found ranging from West Asia and Central Asia to the Indian subcontinent and Southeast Asia. About sixteen subspecies are recognized through its wide range with many island forms ...
(''Saxicola caprata''). In addition mice, lizards, large insects are also taken. In one study in Bangladesh, adults fed mainly on small sparrow sized birds (72%) and ''
Pipistrellus ''Pipistrellus'' is a genus of bats in the family Vespertilionidae and subfamily Vespertilioninae. The name of the genus is derived from the Italian language, Italian word , meaning "bat" (from Latin "bird of evening, bat"). The size of the ge ...
'' bats (28%). They may sometimes
pirate Piracy is an act of robbery or criminal violence by ship or boat-borne attackers upon another ship or a coastal area, typically with the goal of stealing cargo and other valuable goods. Those who conduct acts of piracy are called pirates, v ...
prey obtained by other medium-sized raptors. Prey may sometimes be cached and eaten subsequently. The red-necked falcon drinks water where available during the afternoons. This has been observed both in India and in Africa, where it sometimes visits waterholes. The breeding season in India is January to March. In Zambia, the breeding season begins in August. Pairs may indulge in courtship feeding in which the female feeds the male, an unusual behaviour that has also been noted in captivity. This falcon usually reuses the old tree nest platforms, particularly of
corvid Corvidae is a Cosmopolitan distribution, cosmopolitan family (biology), family of oscine passerine birds that contains the crows, ravens, rook (bird), rooks, jackdaws, jays, magpies, treepies, choughs, and nutcracker (bird), nutcrackers. In coll ...
s, or lays its 3-5 eggs in a nest that it builds on the fork of a tall tree or in the crown of a
palm tree The Arecaceae is a family of perennial flowering plants in the monocot order Arecales. Their growth form can be climbers, shrubs, tree-like and stemless plants, all commonly known as palms. Those having a tree-like form are called palm trees ...
. In Africa, they have been known to reuse the nests of pied crows (''Corvus albus''), African fish eagles (''Halieaetus vocifer'') on ''Acacia'' apart from building their own nest in ''
Borassus ''Borassus'' (palmyra palm) is a genus of five species of fan palms, native to tropical regions of Africa, Asia and Papua New Guinea. Description These massive palms can grow up to high and have robust trunks with distinct leaf scars; in so ...
'' palms. In India, the nest is often placed in a large mango tree (''
Mangifera indica ''Mangifera indica'', commonly known as mango, is a species of flowering plant in the family Anacardiaceae. It is a large fruit tree, capable of growing to a height of . There are two distinct genetic populations in modern mangoesthe "Indian type ...
'') and concealed inside foliage. The nest territory is well-guarded and crows and kites driven away. This falcon has been documented to nests in trees amidst dense human population. The clutch consists of two to four eggs which are incubated only by the female which begins after the last egg of the clutch is laid. The eggs hatch after about 32 to 34 days and the newly hatched young are covered in white down and are brooded by the female for a week. The male brings food which is torn by the female and fed to the chicks. The young fledge in about 35 to 37 days in Africa and up to 48 days in India.


In falconry

The turumti was a favourite among Indian falconers who would fly it especially at the
Indian roller The Indian roller (''Coracias benghalensis'') is a bird of the family Coraciidae. It is long with a wingspan of and weighs . The face and throat are pinkish, the head and back are brown, with blue on the rump and contrasting light and dark blu ...
which would make evasive aerial manoeuvres that entertained the onlookers. These falcons were caught using a
bal-chatri Bal-chatri (/bɑːl tʃʌθri/) are traps designed to catch birds of prey (raptors). The trap essentially consists of a cage baited inside with a conspicuously visible live rodent or small bird, with a series of monofilament nooses attached to th ...
, as they not only captured prey in the air like other falcons, but will also pursue on the ground and thereby could get entangled in the horsehair nooses. Turumta was a name used in Iran for the
merlin Merlin ( cy, Myrddin, kw, Marzhin, br, Merzhin) is a mythical figure prominently featured in the legend of King Arthur and best known as a mage, with several other main roles. His usual depiction, based on an amalgamation of historic and le ...
.


Parasites and diseases

Captive birds can be affected by
Newcastle disease Virulent Newcastle disease (VND), formerly exotic Newcastle disease, is a contagious viral avian disease affecting many domestic and wild bird species; it is transmissible to humans. Though it can infect humans, most cases are non-symptomatic; ...
virus, as well as many parasites including ''Trichomonas'' (in subsp. ''ruficollis'') and nematodes such as ''Cyrnea eurycerca''.


References


External links

* Red-necked falcon
Species text in The Atlas of Southern African Birds
{{Taxonbar, from=Q738666
red-necked falcon The red-necked falcon (''Falco chicquera'') is a bird of prey in the falcon family with two disjunct populations, one in India and the other in Africa. This medium-sized falcon has bluish grey wings and upper body, a chestnut red cap with short c ...
Birds of South Asia Birds of prey of Sub-Saharan Africa
red-necked falcon The red-necked falcon (''Falco chicquera'') is a bird of prey in the falcon family with two disjunct populations, one in India and the other in Africa. This medium-sized falcon has bluish grey wings and upper body, a chestnut red cap with short c ...