Fairy Longhorn Moth
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The Adelidae or fairy longhorn moths are a family of monotrysian moths in the
lepidoptera Lepidoptera ( ) is an order (biology), order of insects that includes butterfly, butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 Family (biology), families and 46 Taxonomic r ...
n infraorder Heteroneura. The family was first described by Charles Théophile Bruand d'Uzelle in 1851. Most species have at least partially metallic patterns coloration and are diurnal, sometimes swarming around the tips of branches with an undulating flight. Others are
crepuscular In zoology, a crepuscular animal is one that is active primarily during the twilight period, being matutinal, vespertine, or both. This is distinguished from diurnal and nocturnal behavior, where an animal is active during the hours of daylig ...
and have a drab coloration. Fairy longhorn moths have a wingspan of 4–28 millimeters, and males often have especially long
antenna Antenna ( antennas or antennae) may refer to: Science and engineering * Antenna (radio), also known as an aerial, a transducer designed to transmit or receive electromagnetic (e.g., TV or radio) waves * Antennae Galaxies, the name of two collid ...
e, 1–3 times as long as the forewing. They are widespread around the world and can be found over much of North America and Eurasia from April to June. About 50 species occur in Europe, of which most widely noted is the
green longhorn The green longhorn (''Adela reaumurella'') is a lepidopteran from the moth family Adelidae, the fairy longhorn moths. Distribution ''Adela reaumurella'' has a palearctic distribution. It occurs in southern North Europe, Western Europe through C ...
(''Adela reaumurella'') which can sometimes reach great abundance; due to climate change its peak flying season is shifting towards spring. In general, they are more plentiful in the Northern Hemisphere, but the family occurs in the Neotropics,
sub-Saharan Africa Sub-Saharan Africa is, geographically, the area and regions of the continent of Africa that lies south of the Sahara. These include West Africa, East Africa, Central Africa, and Southern Africa. Geopolitically, in addition to the List of sov ...
, South-East Asia and
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
too. Adelidae are usually closely restricted to particular host plants, in which the females insert their eggs or just lay among leaf litter, and the caterpillars make a case, completing their development on the ground. Fairy longhorn moths feed in sunshine on
nectar Nectar is a sugar-rich liquid produced by plants in glands called nectaries or nectarines, either within the flowers with which it attracts pollinating animals, or by extrafloral nectaries, which provide a nutrient source to animal mutualists ...
from the flowers of
herbaceous Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition of t ...
(woody) plants.


Systematics

Fairy longhorn moths belong to the
superfamily SUPERFAMILY is a database and search platform of structural and functional annotation for all proteins and genomes. It classifies amino acid sequences into known structural domains, especially into SCOP superfamilies. Domains are functional, str ...
Adeloidea Adeloidea is a superfamily of primitive monotrysian moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are th ...
, one of the
basal Basal or basilar is a term meaning ''base'', ''bottom'', or ''minimum''. Science * Basal (anatomy), an anatomical term of location for features associated with the base of an organism or structure * Basal (medicine), a minimal level that is nec ...
(" monotrysian") branches of the advanced moth infraorder Heteroneura. By
lepidoptera Lepidoptera ( ) is an order (biology), order of insects that includes butterfly, butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 Family (biology), families and 46 Taxonomic r ...
n standards, they are thus still rather primitive micromoths. But like other Heteroneura, they already possess the
apomorph In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have ...
ic sucking proboscis – usually considered a defining feature of Lepidoptera, but the most ancestral moths still live on solid food which they chew.Davis (1999) The Adelidae were previously placed as the subfamily Adelinae within the family Incurvariidae.


Subfamilies

The Adelidae are usually divided into two
subfamilies In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
, but most genera are of uncertain or
basal Basal or basilar is a term meaning ''base'', ''bottom'', or ''minimum''. Science * Basal (anatomy), an anatomical term of location for features associated with the base of an organism or structure * Basal (medicine), a minimal level that is nec ...
relationships. Selected species are also listed: Subfamily Adelinae * ''
Adela Adela may refer to: * ''Adela'', a 1933 Romanian novel by Garabet Ibrăileanu * ''Adela'' (1985 film), a 1985 Romanian film directed by Mircea Veroiu * ''Adela'' (2000 film), a 2000 Argentine thriller film directed and written by Eduardo Mign ...
'' Latreille, 1796 ** ''
Adela croesella ''Adela croesella'' is a moth of the family Adelidae. It is found in most of Europe. The wingspan is 11–14 mm. Head ferruginous mixed with black. Antennae in male 2.5 black, tip white ; in female hardly 1.5, thickened with violet-black sc ...
'' ** ''
Adela reaumurella The green longhorn (''Adela reaumurella'') is a lepidopteran from the moth family Adelidae, the fairy longhorn moths. Distribution ''Adela reaumurella'' has a palearctic distribution. It occurs in southern North Europe, Western Europe through C ...
'' – green longhorn * ''
Cauchas ''Cauchas'' is a genus of the fairy longhorn moth family (Adelidae). Among these, it belongs to subfamily Adelinae. The taxon "Cauchas" Philipp Christoph Zeller 1839 was originally created as a subgenus of the genus ''Adela'' Latreille 1796 but ...
'' Zeller, 1839 ** ''
Cauchas rufimitrella ''Cauchas rufimitrella'' is a diurnal lepidopteran from the family Adelidae, the fairy long horn moths. It is found in almost all of Europe, except Portugal, Ukraine and the southern part of the Balkan Peninsula. The wingspan of the moth range ...
'' * ''
Nemophora ''Nemophora'' is a genus of the Adelidae, fairy longhorn moth family (biology), family (Adelidae). Among these, it belongs to subfamily Adelinae. Selected species * ''Nemophora ahenea'' Stringer, 1930 * ''Nemophora albiciliellus'' (Staudinger, ...
'' Illiger & Hoffmannsegg, 1798 (tentatively placed here) **''
Nemophora degeerella The longhorn moth or yellow-barred long-horn (''Nemophora degeerella'') is a Diurnality, diurnal lepidopteran from the moths family Adelidae (fairy longhorn moths). Distribution and habitat This species is present on most of Europe, but it is fa ...
'' – longhorn moth ** '' Nemophora metallica'' ** ''
Nemophora ochsenheimerella ''Nemophora ochsenheimerella'' is a moth of the family Adelidae. It is a trans- Palearctic species, which is also found in Central Europe. The wingspan is about . The moth flies in one generation from late April to June. Notes #''The flight se ...
'' Subfamily Nematopogoninae * '' Ceromitia'' Zeller, 1852 (tentatively placed here) ** ''
Ceromitia iolampra ''Ceromitia iolampra'' is a moth of the family Adelidae. It is found in the Australian Capital Territory, New South Wales and Victoria (Australia), Victoria. The larvae feed on the flowers of ''Acacia baileyana'' and ''Acacia genistifolia''. La ...
'' * ''
Nematopogon :"Nematopogon" ''was also invalidly established by Agassiz in 1847 as an unjustified emendation for the fungus moth genus '' Nemapogon. __NOTOC__ ''Nematopogon'' is a genus of the fairy longhorn moth family (Adelidae). Among these, it belongs ...
'' '' Incertae sedis'' * ''
Subclemensia ''Subclemensia'' is a genus of moths of the family Incurvariidae Incurvariidae is a family of small primitive monotrysian moths in the order Lepidoptera. There are twelve genera recognised (Davis, 1999). Many species are leaf miners and much i ...
'' * ''
Trichofrons ''Trichofrons'' is a monotypic moth genus of the family Adelidae, the fairy longhorn moths. The genus was erected by Amsel in 1937. Its sole species is ''Trichofrons pantherella'', which was first described by Achille Guenée in 1849. It is found ...
'' The genus ''
Tridentaforma ''Tridentaforma'' is a genus of moths of the Tridentaformidae family, the sole genus of the family. There is only one species in the genus (and the family), ''Tridentaforma fuscoleuca'', which is found in California and Canada. The wingspan ...
'' is sometimes placed among the Adelidae ''incertae sedis'' too; others assign it to the closely related Prodoxidae.


Footnotes


References

* Davis, D. R. (1999): The Monotrysian Heteroneura. ''In:'' Kristensen, N. P. (ed.): ''Handbuch der Zoologie/Handbook of Zoology'' (Volume IV – Arthropoda: Insecta. Part 35: Lepidoptera, Moths and Butterflies 1): 65–90. Walter de Gruyter, Berlin & New York. * Edwards, E. D. (2007): Australian Faunal Directory â€
Adelidae
Version of 28 June 2007. Retrieved 9 May 2010. * Fauna Europaea (FE) (2009)
Adelidae
Version 2.1, 22 December 2009. Retrieved 3 May 2010. * Kuchlein, J. H. & Ellis, W. N. (2004): Climate-induced changes in the microlepidoptera fauna of the Netherlands and the implications for nature conservation. ''Journal of Insect Conservation'' 1(2): 73–80. PDF fulltext
*


External links



* * {{Taxonbar, from=Q927746 Moth families