European wildcat
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The European wildcat (''Felis silvestris'') is a small
wildcat The wildcat is a species complex comprising two small wild cat species: the European wildcat (''Felis silvestris'') and the African wildcat (''F. lybica''). The European wildcat inhabits forests in Europe, Anatolia and the Caucasus, while th ...
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
native to continental
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
,
Scotland Scotland (, ) is a country that is part of the United Kingdom. Covering the northern third of the island of Great Britain, mainland Scotland has a border with England to the southeast and is otherwise surrounded by the Atlantic Ocean to the ...
,
Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a list of transcontinental countries, transcontinental country located mainly on the Anatolia, Anatolian Peninsula in Western Asia, with ...
and the
Caucasus The Caucasus () or Caucasia (), is a region between the Black Sea and the Caspian Sea, mainly comprising Armenia, Azerbaijan, Georgia, and parts of Southern Russia. The Caucasus Mountains, including the Greater Caucasus range, have historically ...
. It inhabits forests from the
Iberian Peninsula The Iberian Peninsula (), ** * Aragonese and Occitan: ''Peninsula Iberica'' ** ** * french: Péninsule Ibérique * mwl, Península Eibérica * eu, Iberiar penintsula also known as Iberia, is a peninsula in southwestern Europe, defi ...
,
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...
,
Central Central is an adjective usually referring to being in the center of some place or (mathematical) object. Central may also refer to: Directions and generalised locations * Central Africa, a region in the centre of Africa continent, also known as ...
and
Eastern Europe Eastern Europe is a subregion of the Europe, European continent. As a largely ambiguous term, it has a wide range of geopolitical, geographical, ethnic, cultural, and socio-economic connotations. The vast majority of the region is covered by Russ ...
to the Caucasus. Its fur is brownish to grey with stripes on the forehead and on the sides and has a bushy tail with a black tip. It reaches a head-to-body length of up to with a long tail, and weighs up to . In
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
and Italy, the European wildcat is predominantly
nocturnal Nocturnality is an animal behavior characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnal meaning the opposite. Nocturnal creatures generally have highly developed sens ...
, but also active in the daytime when undisturbed by human activities. It preys foremost on small mammals such as
lagomorph The lagomorphs are the members of the taxonomic order Lagomorpha, of which there are two living families: the Leporidae (hares and rabbits) and the Ochotonidae (pikas). The name of the order is derived from the Ancient Greek ''lagos'' (λαγ ...
s and
rodent Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are na ...
s, but also on ground-dwelling birds.


Taxonomy

''Felis (catus) silvestris'' was the
scientific name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
proposed in 1778 by Johann von Schreber when he described a wild cat based on texts from the early 18th century and before. In the 19th and 20th centuries, several wildcat
type specimen In biology, a type is a particular wiktionary:en:specimen, specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to a ...
s were described and proposed as
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
, including: * ''Felis silvestris caucasica'' proposed by
Konstantin Satunin Konstantin Alekseevich Satunin (20 May 1863–10 November 1915) was a Russian zoologist who studied and described many mammals found in Russia and Central Asia. Satunin graduated from Moscow University in 1890. From 1893 he worked at a sericultur ...
in 1905 was a skin of a female cat collected near
Borjomi Borjomi ( ka, ბორჯომი) is a resort town in south-central Georgia, 160 km from Tbilisi, with a population of 11,122 (2021). It is one of the municipalities of the Samtskhe–Javakheti region and is situated in the northwestern ...
in
Georgia Georgia most commonly refers to: * Georgia (country), a country in the Caucasus region of Eurasia * Georgia (U.S. state), a state in the Southeast United States Georgia may also refer to: Places Historical states and entities * Related to the ...
. *''Felis grampia'' proposed by Gerrit Smith Miller Jr. in 1907 was a skin and a
skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, the ...
of a male wildcat from
Invermoriston Invermoriston (''Inbhir Mhoireastain'' in Gaelic) () is a small village north of Fort Augustus, Highland, Scotland. The village is on the A82 road, at a junction with the A887. The village's most visited attraction is the Thomas Telford bri ...
in Scotland. Miller revised his classification in 1912, proposing ''Felis silvestris grampia'' after reviewing more wildcat skins from Scotland. *''Felis tartessia'' also proposed by Miller in 1907 was a skin and a skull of a male wildcat from
Jerez de la Frontera Jerez de la Frontera (), or simply Jerez (), is a Spanish city and municipality in the province of Cádiz in the autonomous community of Andalusia, in southwestern Spain, located midway between the Atlantic Ocean and the Cádiz Mountains. , the ...
in southern Spain. The wildcats north of the
Douro The Douro (, , ; es, Duero ; la, Durius) is the highest-flow river of the Iberian Peninsula. It rises near Duruelo de la Sierra in Soria Province, central Spain, meanders south briefly then flows generally west through the north-west part of ...
and
Ebro , name_etymology = , image = Zaragoza shel.JPG , image_size = , image_caption = The Ebro River in Zaragoza , map = SpainEbroBasin.png , map_size = , map_caption = The Ebro ...
Rivers are said to be smaller than in the rest of the region. The disputed "Tartessian" wildcat has kept the same size and proportions as the form that was found in mainland Europe during the
Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...
Ice Ages An ice age is a long period of reduction in the temperature of Earth's surface and atmosphere, resulting in the presence or expansion of continental and polar ice sheets and alpine glaciers. Earth's climate alternates between ice ages and gree ...
. As of 2017, two subspecies are recognised as valid taxa: *''F. s. silvestris'' in continental Europe, Scotland and Sicily *''F. s. caucasica'' in Turkey and the Caucasus.
Zoological specimen A zoological specimen is an animal or part of an animal preserved for scientific use. Various uses are: to verify the identity of a (species), to allow study, increase public knowledge of zoology. Zoological specimens are extremely diverse. Exampl ...
s of cats that originated on
Mediterranean islands The following is a list of islands in the Mediterranean Sea. The two main island countries in the region are Malta and Cyprus, while other countries with islands in the Mediterranean Sea include Italy, France, Greece, Spain, Tunisia, Croatia, ...
are not considered native but introduced, including: *''Felis lybica'' var ''sarda'' proposed in 1885 by
Fernand Lataste Fernand Lataste (1847 - 1934) was a French zoologist and herpetologist born in Cadillac, Gironde. From 1880 to 1884 he collected reptiles and amphibians in North Africa (Algeria, Tunisia and Morocco), publishing ''"Les missions scientifiques d ...
was a skin and a skull of a male cat from
Sarrabus Sarrabus-Gerrei is a sub-region of south-eastern Sardinia, Italy. Sarrabus Traditionally Sarrabus, probably from the Roman-time city of Sarcopos, occupies the area of the communes of Castiadas, Muravera, San Vito and Villaputzu, corresponding to ...
in
Sardinia Sardinia ( ; it, Sardegna, label=Italian, Corsican and Tabarchino ; sc, Sardigna , sdc, Sardhigna; french: Sardaigne; sdn, Saldigna; ca, Sardenya, label=Algherese and Catalan) is the second-largest island in the Mediterranean Sea, after ...
. * ''Felis reyi'' proposed in 1929 by
Louis Lavauden Louis Lavauden (19 June 1881, in Grenoble – 1 September 1935, in Anjou, Isère) was a French zoologist and forester. He was a student at the Institut agronomique et de l'Ecole forestière in Nancy, afterwards conducting zoological studi ...
who described a skin and a skull of a specimen from
Biguglia Biguglia () is a commune in the Haute-Corse department of France on the island of Corsica. It is near the town of Bastia. Population Sport Biguglia is the home of Championnat de France Amateurs 2 club, ÉF Bastia. See also * Communes ...
. * ''F. s. cretensis'' proposed in 1953 by
Theodor Haltenorth Dr. Theodor Haltenorth (May 18, 1910 – January 30, 1981) was a German mammalogist. He worked mainly in Munich, Bavaria, Germany, and was a key figure in the Quagga Project. He also taxonomised the Cretan wildcat The Cretan wildcat is a me ...
who described two cat skins that were purchased in a bazaar in
Chania Chania ( el, Χανιά ; vec, La Canea), also spelled Hania, is a city in Greece and the capital of the Chania regional unit. It lies along the north west coast of the island Crete, about west of Rethymno and west of Heraklion. The muni ...
.


Phylogeny

Phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
analysis of the
nuclear DNA Nuclear DNA (nDNA), or nuclear deoxyribonucleic acid, is the DNA contained within each cell nucleus of a eukaryotic organism. It encodes for the majority of the genome in eukaryotes, with mitochondrial DNA and plastid DNA coding for the rest. It ...
in tissue samples from all
Felidae Felidae () is the family of mammals in the order Carnivora colloquially referred to as cats, and constitutes a clade. A member of this family is also called a felid (). The term "cat" refers both to felids in general and specifically to the ...
species revealed that the
evolutionary radiation An evolutionary radiation is an increase in taxonomic diversity that is caused by elevated rates of speciation, that may or may not be associated with an increase in morphological disparity. Radiations may affect one clade or many, and be rapid ...
of the Felidae began in Asia in the
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
around . Analysis of
mitochondrial DNA Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial D ...
of all Felidae species indicates a radiation at around . The European wildcat is part of an
evolution Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variation ...
ary
lineage Lineage may refer to: Science * Lineage (anthropology), a group that can demonstrate its common descent from an apical ancestor or a direct line of descent from an ancestor * Lineage (evolution), a temporal sequence of individuals, populati ...
that is estimated to have genetically diverged from the
common ancestor Common descent is a concept in evolutionary biology applicable when one species is the ancestor of two or more species later in time. All living beings are in fact descendants of a unique ancestor commonly referred to as the last universal comm ...
of the ''Felis'' species around , based on analysis of their nuclear DNA. Analysis of their mitochondrial DNA indicates a genetic divergence from ''Felis'' at around . Both models agree in the
jungle cat The jungle cat (''Felis chaus''), also called reed cat, swamp cat and jungle lynx, is a medium-sized cat native to the Middle East, the Caucasus, South and Southeast Asia and southern China. It inhabits foremost wetlands like swamps, littora ...
(''F. chaus'') having been the first ''Felis'' species that diverged, followed by the
black-footed cat The black-footed cat (''Felis nigripes''), also called the small-spotted cat, is the smallest wild cat in Africa, having a head-and-body length of . Despite its name, only the soles of its feet are black or dark blackish brown. With its bold s ...
(''F. nigripes''), the
sand cat The sand cat (''Felis margarita'') is a small wild cat that inhabits sandy and stony deserts far from water sources. With its sandy to light grey fur, it is well camouflaged in a desert environment. Its head-and-body length ranges from with a ...
(''F. margarita''), the
African wildcat The African wildcat (''Felis lybica'') is a small wildcat species native to Africa, West and Central Asia up to Rajasthan in India and Xinjiang in China. It has been listed as Least Concern on the IUCN Red List in 2022. In Cyprus, an African wil ...
(''F. lybica'') and then the European wildcat.
Fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
remains of small wild cats found in Europe indicate that the European wildcat probably descended from ''
Felis lunensis ''Felis lunensis,'' or the Martelli's cat is an extinct felid of the subfamily Felinae. Evolution and taxonomy Around 12 million years ago, the genus '' Felis'' appeared and eventually gave rise to many of the modern small cats. ''Felis lun ...
'' in the
Villafranchian Villafranchian age ( ) is a period of geologic time (3.5–1.0 Ma) spanning the Late Pliocene and Early Pleistocene used more specifically with European Land Mammal Ages. Named by Italian geologist Lorenzo Pareto for a sequence of terrestrial s ...
more than , a transition that was completed by the Holstein interglacial about 340,000 to 325,000 years ago.


Characteristics

The European wildcat's fur varies in colour from brownish to grey with paler contour hairs. It has five stripes on the forehead, which are broken up into small spots. A dark stripe behind the shoulders expands into a spinal stripe running up to the base of the tail. On the sides, it has irregular dark stripes, which break up on the hind legs, thus forming a blotched pattern. Its tail is bushy with two to three black, transverse rings and rounded at the black tip. The top of the head and the forehead bear four well-developed dark bands that split into small spots. Two short and narrow stripes are usually present in the shoulder region, in front of the dorsal band. Some individuals have a few light spots on the throat, between the forelegs, or in the inguinal region. The dorsal surface of the neck and head are the same colour as that of the trunk, but is lighter grey around the eyes, lips, cheeks, and chin. A slight ochreous shade is visible on the undersides of the flanks. A black and narrow dorsal band starts on the shoulders, and runs along the back up to the base of the tail. In some animals, the summer coat is ashen coloured. The patterns on the head and neck are as well-developed as those on the tail, though the patterns on the flanks are almost imperceptible.
Guard hair Guard hair or overhair is the outer layer of hair of most mammals, which overlay the fur. Guard hairs are long and coarse and protect the rest of the pelage (fur) from abrasion and frequently from moisture. They are visible on the surface of the fu ...
s measure , the tip hairs , and the underfur . Corresponding measurements in the summer are , , and . Large males in Spain reach in length, with a long tail, and weigh up to . They also have a less diffuse stripe pattern, proportionally larger teeth, and feed more often on
rabbit Rabbits, also known as bunnies or bunny rabbits, are small mammals in the family Leporidae (which also contains the hares) of the order Lagomorpha (which also contains the pikas). ''Oryctolagus cuniculus'' includes the European rabbit speci ...
s than the wildcats north of the Douro-Ebro, which are more dependent on small
rodent Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are na ...
s. The European wildcat is on average bigger and stouter than the domestic
cat The cat (''Felis catus'') is a domestic species of small carnivorous mammal. It is the only domesticated species in the family Felidae and is commonly referred to as the domestic cat or house cat to distinguish it from the wild members of ...
, has longer
fur Fur is a thick growth of hair that covers the skin of mammals. It consists of a combination of oily guard hair on top and thick underfur beneath. The guard hair keeps moisture from reaching the skin; the underfur acts as an insulating blanket t ...
and a shorter non-tapering bushy tail. It has striped fur and a dark dorsal band. Males average a weight of up to , and females . Their weight fluctuates seasonally up to . European wildcats have proportionately shorter cheek tooth rows with smaller teeth, but a broader muzzle than African wildcats. Since European wildcats and domestic cats opportunistically
interbreed In biology, a hybrid is the offspring resulting from combining the qualities of two organisms of different breeds, varieties, species or genera through sexual reproduction. Hybrids are not always intermediates between their parents (such as in ...
, it is difficult to distinguish wildcats and striped hybrids correctly on the basis of only morphological characteristics.


Distribution and habitat

The European wildcat lives primarily in
broad-leaved A broad-leaved, broad-leaf, or broadleaf tree is any tree within the diverse botanical group of angiosperms that has flat leaves and produces seeds inside of fruits. It is one of two general types of trees, the other being a conifer, a tree with ne ...
and mixed forests. It avoids intensively cultivated areas and settlements. The northernmost population lives in northern and eastern Scotland. It has been
extirpated Local extinction, also known as extirpation, refers to a species (or other taxon) of plant or animal that ceases to exist in a chosen geographic area of study, though it still exists elsewhere. Local extinctions are contrasted with global extinct ...
in England and Wales. There are two disconnected populations in France. The one in the
Ardennes The Ardennes (french: Ardenne ; nl, Ardennen ; german: Ardennen; wa, Årdene ; lb, Ardennen ), also known as the Ardennes Forest or Forest of Ardennes, is a region of extensive forests, rough terrain, rolling hills and ridges primarily in Be ...
in the country's north-east extends to
Luxembourg Luxembourg ( ; lb, Lëtzebuerg ; french: link=no, Luxembourg; german: link=no, Luxemburg), officially the Grand Duchy of Luxembourg, ; french: link=no, Grand-Duché de Luxembourg ; german: link=no, Großherzogtum Luxemburg is a small lan ...
,
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
and
Belgium Belgium, ; french: Belgique ; german: Belgien officially the Kingdom of Belgium, is a country in Northwestern Europe. The country is bordered by the Netherlands to the north, Germany to the east, Luxembourg to the southeast, France to th ...
. The other in southern France may be connected via the
Pyrenees The Pyrenees (; es, Pirineos ; french: Pyrénées ; ca, Pirineu ; eu, Pirinioak ; oc, Pirenèus ; an, Pirineus) is a mountain range straddling the border of France and Spain. It extends nearly from its union with the Cantabrian Mountains to C ...
to populations in Spain and Portugal. In the Netherlands, European wildcats were recorded in 1999 near
Nijmegen Nijmegen (;; Spanish and it, Nimega. Nijmeegs: ''Nimwèège'' ) is the largest city in the Dutch province of Gelderland and tenth largest of the Netherlands as a whole, located on the Waal river close to the German border. It is about 6 ...
and in 2004 in
North Brabant North Brabant ( nl, Noord-Brabant ; Brabantian: ; ), also unofficially called Brabant, is a province in the south of the Netherlands. It borders the provinces of South Holland and Gelderland to the north, Limburg to the east, Zeeland to the we ...
; these individuals had possibly dispersed from Germany. In Germany, the
Rhine ), Surselva, Graubünden, Switzerland , source1_coordinates= , source1_elevation = , source2 = Rein Posteriur/Hinterrhein , source2_location = Paradies Glacier, Graubünden, Switzerland , source2_coordinates= , so ...
is a major barrier between the population in
Eifel The Eifel (; lb, Äifel, ) is a low mountain range in western Germany and eastern Belgium. It occupies parts of southwestern North Rhine-Westphalia, northwestern Rhineland-Palatinate and the southern area of the German-speaking Community of ...
and
Hunsrück The Hunsrück () is a long, triangular, pronounced upland in Rhineland-Palatinate, Germany. It is bounded by the valleys of the Moselle-Saar (north-to-west), the Nahe (south), and the Rhine (east). It is continued by the Taunus mountains, past ...
mountains west of the river and populations east of the river, where a six-lane highway hampers dispersal. In Switzerland, European wildcats are present in the
Jura Mountains The Jura Mountains ( , , , ; french: Massif du Jura; german: Juragebirge; it, Massiccio del Giura, rm, Montagnas da Jura) are a sub-alpine mountain range a short distance north of the Western Alps and mainly demarcate a long part of the Frenc ...
. Three fragmented populations in Italy comprise one in the country's central and southern part, one in the eastern
Alps The Alps () ; german: Alpen ; it, Alpi ; rm, Alps ; sl, Alpe . are the highest and most extensive mountain range system that lies entirely in Europe, stretching approximately across seven Alpine countries (from west to east): France, Sw ...
that may be connected to populations in Slovenia and Croatia. The Sicilian population is the only Mediterranean insular population that has not been introduced. The population in the Polish
Carpathian Mountains The Carpathian Mountains or Carpathians () are a range of mountains forming an arc across Central Europe. Roughly long, it is the third-longest European mountain range after the Urals at and the Scandinavian Mountains at . The range stretches ...
extends to northern Slovakia and western Ukraine.


Behaviour and ecology

In France and Italy, the European wildcat is active foremost at night; in undisturbed sites, it is also active by day. In Sicily, an individual was photographed in 2009 and again in 2018 at about the same location. It was probably at least 10 years old at the time of recapture.


Hunting and diet

In Western Europe, the wildcat feeds on
hamster Hamsters are rodents (order Rodentia) belonging to the subfamily Cricetinae, which contains 19 species classified in seven genera.Fox, Sue. 2006. ''Hamsters''. T.F.H. Publications Inc. They have become established as popular small pets. The b ...
s,
brown rat The brown rat (''Rattus norvegicus''), also known as the common rat, street rat, sewer rat, wharf rat, Hanover rat, Norway rat, Norwegian rat and Parisian rat, is a widespread species of common rat. One of the largest muroids, it is a brown o ...
s, dormice, water voles, voles, and wood mice. From time to time, it also preys on small carnivores like martens,
European polecat The European polecat (''Mustela putorius''), also known as the common polecat, black polecat, or forest polecat, is a species of mustelid native to western Eurasia and North Africa. It is of a generally dark brown colour, with a pale underbelly ...
,
stoat The stoat (''Mustela erminea''), also known as the Eurasian ermine, Beringian ermine and ermine, is a mustelid native to Eurasia and the northern portions of North America. Because of its wide circumpolar distribution, it is listed as Least Conc ...
, and
least weasel The least weasel (''Mustela nivalis''), little weasel, common weasel, or simply weasel is the smallest member of the genus '' Mustela,'' family Mustelidae and order Carnivora. It is native to Eurasia, North America and North Africa, and has bee ...
(''Mustela nivalis''), as well as fawns of
red deer The red deer (''Cervus elaphus'') is one of the largest deer species. A male red deer is called a stag or hart, and a female is called a hind. The red deer inhabits most of Europe, the Caucasus Mountains region, Anatolia, Iran, and parts of wes ...
(''Cervus elaphus''),
roe deer The roe deer (''Capreolus capreolus''), also known as the roe, western roe deer, or European roe, is a species of deer. The male of the species is sometimes referred to as a roebuck. The roe is a small deer, reddish and grey-brown, and well-adapt ...
(''Capreolus capreolus''), and
chamois The chamois (''Rupicapra rupicapra'') or Alpine chamois is a species of goat-antelope native to mountains in Europe, from west to east, including the Alps, the Dinarides, the Tatra and the Carpathian Mountains, the Balkan Mountains, the Ril ...
(''Rupicapra rupicapra''). In the
Carpathians The Carpathian Mountains or Carpathians () are a range of mountains forming an arc across Central Europe. Roughly long, it is the third-longest European mountain range after the Ural Mountains, Urals at and the Scandinavian Mountains at . The ...
, the wildcat feeds primarily on
yellow-necked mouse The yellow-necked mouse (''Apodemus flavicollis''), also called yellow-necked field mouse, yellow-necked wood mouse, and South China field mouse, is closely related to the wood mouse, with which it was long confused. It was only recognised as a ...
(''Apodemus flavicollis''),
northern red-backed vole The northern red-backed vole (''Myodes rutilus'') is a small slender vole found in Alaska, northern Canada, Scandinavia and northern Russia.Banfield, A. W. F. 1974. The mammals of Canada. Toronto, University of Toronto Press. Description They h ...
(''Myodes rutilus''), Tatra pine vole (''Microtus tatricus''), and occasionally also
European hare The European hare (''Lepus europaeus''), also known as the brown hare, is a species of hare native to Europe and parts of Asia. It is among the largest hare species and is adapted to temperate, open country. Hares are herbivorous and feed mainly ...
(''Lepus europaeus''). In
Transcarpathia Transcarpathia may refer to: Place * relative term, designating any region beyond the Carpathians (lat. ''trans-'' / beyond, over), depending on a point of observation * Romanian Transcarpathia, designation for Romanian regions on the inner or ...
, the wildcat's diet consists of mouse-like rodents,
galliformes Galliformes is an order of heavy-bodied ground-feeding birds that includes turkeys, chickens, quail, and other landfowl. Gallinaceous birds, as they are called, are important in their ecosystems as seed dispersers and predators, and are often ...
, and
squirrel Squirrels are members of the family Sciuridae, a family that includes small or medium-size rodents. The squirrel family includes tree squirrels, ground squirrels (including chipmunks and prairie dogs, among others), and flying squirrels. Squ ...
s. In the Dnestr swamps, it preys on ''
Microtus ''Microtus'' is a genus of voles found in North America, Europe and northern Asia. The genus name refers to the small ears of these animals. About 62 species are placed in the genus. They are stout rodents with short ears, legs and tails. They ea ...
'', water voles, and birds, while those living in the
Prut The Prut (also spelled in English as Pruth; , uk, Прут) is a long river in Eastern Europe. It is a left tributary of the Danube. In part of its course it forms Romania's border with Moldova and Ukraine. Characteristics The Prut originates ...
swamps primarily target water vole, brown rat, and
muskrat The muskrat (''Ondatra zibethicus'') is a medium-sized semiaquatic rodent native to North America and an introduced species in parts of Europe, Asia, and South America. The muskrat is found in wetlands over a wide range of climates and habitat ...
(''Ondatra zibethicus''). Birds taken by Prut wildcats include
warbler Various Passeriformes (perching birds) are commonly referred to as warblers. They are not necessarily closely related to one another, but share some characteristics, such as being fairly small, vocal, and insectivorous. Sylvioid warblers Th ...
s,
ferruginous duck The ferruginous duck (''Aythya nyroca''), also known as ferruginous pochard, common white-eye or white-eyed pochard, is a medium-sized diving duck from Eurosiberia. The scientific name is derived from Greek '' aithuia'' an unidentified seabir ...
,
Eurasian coot The Eurasian coot (''Fulica atra''), also known as the common coot, or Australian coot, is a member of the rail and crake bird family, the Rallidae. It is found in Europe, Asia, Australia, New Zealand and parts of North Africa. It has a slaty-bla ...
,
spotted crake The spotted crake (''Porzana porzana'') is a small waterbird of the family Rallidae. The scientific name is derived from Venetian terms for small rails. The spotted crake's breeding habitat is marshes and sedge beds across temperate Europe into ...
, and gadwall. In
Moldavia Moldavia ( ro, Moldova, or , literally "The Country of Moldavia"; in Romanian Cyrillic: or ; chu, Землѧ Молдавскаѧ; el, Ἡγεμονία τῆς Μολδαβίας) is a historical region and former principality in Centr ...
, the wildcat's winter diet consists primarily of rodents, while it preys on birds,
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...
, and
crayfish Crayfish are freshwater crustaceans belonging to the clade Astacidea, which also contains lobsters. In some locations, they are also known as crawfish, craydids, crawdaddies, crawdads, freshwater lobsters, mountain lobsters, rock lobsters, mu ...
in summer. Brown rats and water voles, as well as muskrats and
waterfowl Anseriformes is an order of birds also known as waterfowl that comprises about 180 living species of birds in three families: Anhimidae (three species of screamers), Anseranatidae (the magpie goose), and Anatidae, the largest family, which in ...
are the main sources of food for wildcats in the
Kuban River The Kuban; Circassian: Псыжъ, ''Psyẑ'' or Псыжь, ''Psyź'' ; abq, Къвбина, ''Q̇vbina'' ; Karachay–Balkar: Къобан, ''Qoban''; Nogai: Кобан, ''Qoban'') is a river in Russia that flows through the Western Cauca ...
delta. Wildcats in the northern Caucasus feed on mouse-like rodents and edible dormice, as well as birds, young chamois and roe deer on rare occasions. Wildcats on the
Black Sea The Black Sea is a marginal mediterranean sea of the Atlantic Ocean lying between Europe and Asia, east of the Balkans, south of the East European Plain, west of the Caucasus, and north of Anatolia. It is bounded by Bulgaria, Georgia, Roma ...
coast are thought to feed on small birds, shrews, and hares. On one occasion, the feathers of a
white-tailed eagle The white-tailed eagle (''Haliaeetus albicilla'') is a very large species of sea eagle widely distributed across temperate Eurasia. Like all eagles, it is a member of the family Accipitridae (or accipitrids) which includes other diurnal raptors ...
and the skull of a
kid Kid, Kids, KIDS, and K.I.D.S. may refer to: Common meanings * Colloquial term for a child or other young person ** Also for a parent's offspring regardless of age * Engage in joking * Young goats * The goat meat of young goats * Kidskin, lea ...
were found at a den site. In
Transcaucasia The South Caucasus, also known as Transcaucasia or the Transcaucasus, is a geographical region on the border of Eastern Europe and Western Asia, straddling the southern Caucasus Mountains. The South Caucasus roughly corresponds to modern Arme ...
, the wildcat's diet consists of
gerbil The Mongolian gerbil or Mongolian jird (''Meriones unguiculatus'') is a small rodent belonging to the subfamily Gerbillinae. Their body size is typically , with a tail, and body weight , with adult males larger than females. The animal is use ...
s, voles, birds, and
reptile Reptiles, as most commonly defined are the animals in the class Reptilia ( ), a paraphyletic grouping comprising all sauropsids except birds. Living reptiles comprise turtles, crocodilians, squamates (lizards and snakes) and rhynchocephalians ( ...
s in the summer, and birds, mouse-like rodents, and hares in winter. The Scottish wildcat mainly preys on
European rabbit The European rabbit (''Oryctolagus cuniculus'') or coney is a species of rabbit native to the Iberian Peninsula (including Spain, Portugal, and southwestern France), western France, and the northern Atlas Mountains in northwest Africa. It has ...
(''Oryctolagus cuniculus''),
field vole The short-tailed field vole, short-tailed vole, or simply field vole (''Microtus agrestis'') is a grey-brown vole, around 10 cm in length, with a short tail. It is one of the most common mammals in Europe, with a range extending from the Atl ...
(''Microtus agrestis''), bank vole (''Myodes glareolus''),
wood mouse The wood mouse (''Apodemus sylvaticus'') is a murid rodent native to Europe and northwestern Africa. It is closely related to the yellow-necked mouse (''Apodemus flavicollis'') but differs in that it has no band of yellow fur around the neck, ha ...
(''Apodemus sylvaticus''), and birds.


Threats

In most European countries, European wildcats have become rare. Although legally protected, they are still shot by people mistaking them for feral cats. In the
Scottish Highlands The Highlands ( sco, the Hielands; gd, a’ Ghàidhealtachd , 'the place of the Gaels') is a historical region of Scotland. Culturally, the Highlands and the Lowlands diverged from the Late Middle Ages into the modern period, when Lowland Sco ...
, where approximately 400 were thought to remain in the wild in 2004,
interbreed In biology, a hybrid is the offspring resulting from combining the qualities of two organisms of different breeds, varieties, species or genera through sexual reproduction. Hybrids are not always intermediates between their parents (such as in ...
ing with feral cats is a significant threat to the wild population's distinctiveness. The population in Portugal and Spain is also threatened by interbreeding with feral cats and loss of habitat. The extent of hybridization is low in Germany, Italy and Luxembourg. In the 1990s, the easternmost population in Ukraine, Moldova, and the
Caucasus The Caucasus () or Caucasia (), is a region between the Black Sea and the Caspian Sea, mainly comprising Armenia, Azerbaijan, Georgia, and parts of Southern Russia. The Caucasus Mountains, including the Greater Caucasus range, have historically ...
was threatened by destruction of broad-leaved forests, entailing a reduction of their range. Only small numbers occur in protected areas.


Conservation efforts

The European wildcat is protected in most European range countries. It is listed in
CITES Appendix II CITES (shorter name for the Convention on International Trade in Endangered Species of Wild Fauna and Flora, also known as the Washington Convention) is a multilateral treaty to protect endangered plants and animals from the threats of interna ...
, in Appendix II of the
Berne Convention on the Conservation of European Wildlife and Natural Habitats The Bern Convention on the Conservation of European Wildlife and Natural Habitats, also known as the Bern Convention (or Berne Convention), is a binding international legal instrument in the field of Nature Conservation, it covers the natural h ...
and in the
European Union The European Union (EU) is a supranational political and economic union of member states that are located primarily in Europe. The union has a total area of and an estimated total population of about 447million. The EU has often been des ...
's Habitats and Species Directive.


Germany

In 2004, the Friends of the Earth Germany initiated the project "Safety Net for the European Wildcat". This project aimed at relinking Germany's forests by planting bushes and trees between areas inhabited by and suitable for European wildcat, and which are larger than . They developed the "Wildcat Routing Map", a map depicting the long network of corridors. An Action Plan for the Protection of the European Wildcat in Germany was developed in 2009, aiming at doubling the area inhabited by European wildcat and linking populations within Germany and with neighbouring countries until 2019.


Scotland

In 2013, the Scottish Wildcat Conservation Action Group developed the Scottish Wildcat Conservation Action Plan. With this plan, the group set national action priorities and defined responsibilities of agencies and funding priorities for conservation efforts between 2013 and 2019. Its implementation is coordinated by
Scottish Natural Heritage NatureScot ( gd, NàdarAlba), which was formerly known as Scottish Natural Heritage, is an executive non-departmental public body of the Scottish Government responsible for the country's natural heritage, especially its natural, genetic and ...
. However, the population has been deemed no longer viable.


In captivity

The European wildcat has the reputation for being effectively impossible to raise as a pet. Naturalist Frances Pitt wrote "there was a time when I did not believe this ... my optimism was daunted" by trying to keep a wildcat she named Beelzebina. In England, conservationists plan to start a captive breeding programme in 2019 with the aim to reintroduce cats into the wild by 2022.


References


External links

* * * {{Authority control Felis Wildcats Mammals described in 1777 Fauna of Albania Felids of Europe