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''Eucryphia'' is a small
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomencla ...
of trees and large shrubs native to the south temperate regions of South America and coastal eastern Australia, mainly Tasmania. Sometimes placed in a family of their own, the Eucryphiaceae, more recent classifications place them in the
Cunoniaceae Cunoniaceae is a family of 27 genera and about 335 species of woody plants in the order Oxalidales, mostly found in the tropical and wet temperate regions of the Southern Hemisphere. The greatest diversity of genera are in Australia and Tasmania ...
. There are seven species, two in South America and five in Australia, and several named hybrids.


Description

They are mostly
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, which ...
though one species (''E. glutinosa'') is usually
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, afte ...
. The leaves are opposite, and either simple or pinnate with 3-13 leaflets. The flowers are produced in late summer or autumn, are showy and sweetly scented, 3–6 cm diameter, with four creamy-white
petal Petals are modified leaves that surround the reproductive parts of flowers. They are often brightly colored or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''corolla''. Petals are usually ...
s, and numerous
stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...
s and styles. The fruit is a woody capsule 1-1.5 cm long containing several
seed A seed is an Plant embryogenesis, embryonic plant enclosed in a testa (botany), protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, includ ...
s, and maturing in 12–15 months.


Etymology

The generic name ''Eucryphia'' is composed of two parts, namely ''eu-'' and ''-cryphia''. The Greek ευ-κρυφαιος means well-covered and refers to the foliage, which is clustered towards the apex of braches.


Species


Extant species

*''
Eucryphia cordifolia ''Eucryphia cordifolia'', the ulmo, is a species of tree in the family Cunoniaceae. It is found in Chile and Argentina. It is threatened by logging and habitat loss. The natural habitat is along the Andes Range from 38 to 43°S, and up to 700 m ...
'' Cav.
Valdivian temperate rain forests The Valdivian temperate forests (NT0404) is an ecoregion on the west coast of southern South America, in Chile and Argentina. It is part of the Neotropical realm. The forests are named after the city of Valdivia. The Valdivian temperate rainfores ...
of southern
Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a ...
and
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
. A tree growing up to 40 m tall; leaves simple, crenate to serrated, evergreen, 3–7 cm long. Its wood is used for construction, furniture and lumber, and its nectar for
honey Honey is a sweet and viscous substance made by several bees, the best-known of which are honey bees. Honey is made and stored to nourish bee colonies. Bees produce honey by gathering and then refining the sugary secretions of plants (primaril ...
. *'' Eucryphia glutinosa'' (
Poepp. Eduard Friedrich Poeppig (16 July 1798 – 4 September 1868) was a German botanist, zoologist and explorer. Biography Poeppig was born in Plauen, Saxony. He studied medicine and natural history at the University of Leipzig, graduating with a ...
&
Endl. Stephan Ladislaus Endlicher also known as Endlicher István László (24 June 1804, Bratislava (Pozsony) – 28 March 1849, Vienna) was an Austrian botanist, numismatist and Sinologist. He was a director of the Botanical Garden of Vienna. Bi ...
) Baill.
Valdivian temperate rain forests The Valdivian temperate forests (NT0404) is an ecoregion on the west coast of southern South America, in Chile and Argentina. It is part of the Neotropical realm. The forests are named after the city of Valdivia. The Valdivian temperate rainfores ...
of southern
Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a ...
. A large shrub to 7 m tall; leaves pinnate, with 3-5 leaflets, serrated, deciduous or semi-evergreen, 3–6 cm long. *''
Eucryphia jinksii ''Eucryphia jinksii'', the Springbrook leatherwood, is a species of rare rainforest trees found in Queensland and New South Wales, Australia, of the plant family Cunoniaceae. They grow naturally to 30m tall yet were discovered as a new species ...
'' P.I.Forst. – subtropical
New South Wales ) , nickname = , image_map = New South Wales in Australia.svg , map_caption = Location of New South Wales in AustraliaCoordinates: , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , e ...
&
Queensland ) , nickname = Sunshine State , image_map = Queensland in Australia.svg , map_caption = Location of Queensland in Australia , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , established_ ...
mountain rainforests endemic, Australia. A tree to 25 m tall; leaves simple or with 3 leaflets, entire, evergreen, 5–10 cm long (discovered 1994). *''
Eucryphia lucida ''Eucryphia lucida'', the leatherwood, is a species of tree or large shrub endemic to forests of western Tasmania, Australia. An attractive plant used in both horticulture and apiculture, it was promoted by the Tasmanian Branch of the then SG ...
'' ( Labill.) Baill.
Tasmania ) , nickname = , image_map = Tasmania in Australia.svg , map_caption = Location of Tasmania in AustraliaCoordinates: , subdivision_type = Country , subdi ...
Australia. A tree to 20–30 m tall; leaves simple, entire, evergreen, 3–7 cm long. Common name is Tasmanian leatherwood and it is a major commercial source of
monofloral honey Monofloral honey is a type of honey which has a distinctive flavor or other attribute due to its being predominantly from the nectar of a single plant species. It is stored and labeled separately so as to command a premium price. While there may ...
. *''
Eucryphia milliganii ''Eucryphia milliganii'', also known as the dwarf leatherwood, is a shrub or small tree endemic to areas of Tasmania. It grows in western and southern Tasmania, where it is most commonly found in alpine and sub-alpine heath areas. It is the sis ...
'' Hook.f. (syn. ''E. lucida'' var. ''milliganii''). –
Tasmania ) , nickname = , image_map = Tasmania in Australia.svg , map_caption = Location of Tasmania in AustraliaCoordinates: , subdivision_type = Country , subdi ...
Australia. A tree to 6 m tall or more often a shrub; leaves simple, entire, evergreen, 1.5–3 cm long. *''
Eucryphia moorei ''Eucryphia moorei'', commonly known as pinkwood, plumwood, or eastern leatherwood is a tree found in southeastern New South Wales, Australia. It also occurs just over the border at the Howe Range in Victoria. Pinkwood is the dominant tree speci ...
'' F.Muell.
New South Wales ) , nickname = , image_map = New South Wales in Australia.svg , map_caption = Location of New South Wales in AustraliaCoordinates: , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , e ...
,
Victoria Victoria most commonly refers to: * Victoria (Australia), a state of the Commonwealth of Australia * Victoria, British Columbia, provincial capital of British Columbia, Canada * Victoria (mythology), Roman goddess of Victory * Victoria, Seyche ...
temperate rainforests, Australia. Also known as pinkwood or plumwood, is a tree to 30 m tall; leaves pinnate, with 7-15 leaflets, entire, evergreen, 6–12 cm long. *'' Eucryphia wilkiei'' B.HylandQueensland wet tropics mountain top cloud forests endemic, Australia, is a large shrub. Discovered in Jan. 1970 by Jack (John H.) Wilkie.


Extinct species

*'' Eucryphia falcata'' (Late
Palaeocene The Paleocene, ( ) or Palaeocene, is a geological epoch that lasted from about 66 to 56 million years ago (mya). It is the first epoch of the Paleogene Period in the modern Cenozoic Era. The name is a combination of the Ancient Greek ''palaió ...
, Lake Bungarby) *'' Eucryphia microstoma'' (Early
Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', "daw ...
, Regatta Point) *'' Eucryphia aberensis'' (Middle to Late
Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', "daw ...
, Loch Aber).


Natural hybrids

There are two known natural hybrids, although additional, artificial hybrids have been created. However, these are not naturally occurring. *'' Eucryphia × hybrida'' J.Bausch - natural hybrid of ''E. lucida'' and ''E. milliganii'' native to Tasmania *'' Eucryphia × nymansensis'' J.Bausch - natural hybrid of ''E. cordifolia '' and ''E. glutinosa'' native to Chile


Artificial hybrids and cultivars

(those marked have gained the
Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr (N ...
's
Award of Garden Merit The Award of Garden Merit (AGM) is a long-established annual award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. History The Award of Garden Merit ...
) *''E.'' × ''intermedia'' (''E. glutinosa'' × ''E. lucida'') **''E.'' × ''intermedia'' 'Rostrevor' *''E.'' × ''hillieri'' (''E. moorei'' × ''E. lucida''): Developed from a self-sown seedling at the nursery of Hillier & Sons, Chandlers Ford in Hampshire around 1953. *''E.'' × ''nymansensis'' (''E. cordifolia'' × ''E. glutinosa'') **''E.'' × ''nymansensis'' 'Nymansay' originated from Lt. Col. Leonard C.R. Messel's garden at Nymans, Sussex (1913) **''E.'' × ''nymansensis'' 'Nymans Silver' is a sport discovered at Nymans, Sussex in 2005. It is a variegated form with serrated oval leaves that are outlined in creamy white. It was discovered growing on a ''E.'' × ''nymansensis'' within the garden. It is considered to be faster growing and more columnar in nature. The name 'Nymans Silver' was proposed by Philip Holmes, Deputy Head Gardener at Nymans.

*''E.'' 'Penwith' (''E. cordifolia'' × ''E. lucida'') *''E.'' × ''hybrida'' (''E. glutinosa'' × ''E. lucida'')


Uses

The species and their hybrids are attractive small trees for
garden A garden is a planned space, usually outdoors, set aside for the cultivation, display, and enjoyment of plants and other forms of nature. The single feature identifying even the wildest wild garden is ''control''. The garden can incorporate both ...
s, typically with a slender conic crown when young, though widening with age. They are valued for their conspicuous scented flowers, produced in late summer and autumn when few or no other trees are in flower. Cultivation is restricted to areas with mild winters, cool summers and good rainfall; away from their native areas, this restricts them to the Atlantic coastal regions of Europe, the
Pacific Northwest The Pacific Northwest (sometimes Cascadia, or simply abbreviated as PNW) is a geographic region in western North America bounded by its coastal waters of the Pacific Ocean to the west and, loosely, by the Rocky Mountains to the east. Though ...
of
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the C ...
, and
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island country ...
.


Honey

The nectar of two of the species provides an important sources of honey. ''Eucryphia lucida'' from Tasmania is the main source of a very distinctively flavoured honey known as Leatherwood (the common name for the species). Some of this honey may come from the other Tasmanian species, ''E. milliganii''. In Chile, Ulmo honey (again after the local species name) comes from ''E. cordifolia''. Leatherwood honey and Ulmo honey are very similar in flavour, even though the two species have probably been separated for more than 45 million years.


References


External links


''Eucryphia'' occurrence data
from
GBIF The Global Biodiversity Information Facility (GBIF) is an international organisation that focuses on making scientific data on biodiversity available via the Internet using web services. The data are provided by many institutions from around th ...

''Eucryphia'' in Encyclopedia of the Chilean Flora
* Pictures o

an

in Chile. {{Taxonbar, from=Q639384 Oxalidales genera Flora of Argentina Flora of Chile Oxalidales of Australia Flora of Tasmania Taxa named by Antonio José Cavanilles