Eotitanosuchus Olsoni
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''Eotitanosuchus'' ("dawn giant crocodile") is an extinct genus of biarmosuchian therapsids whose fossils were found in the town of
Ochyor Ochyor (russian: Очёр), alternatively spelled Ocher, is a town and the administrative center of Ochyorsky District in Perm Krai, Russia, located on the Ochyor River (Kama's tributary), west of Perm, the administrative center of the krai. Po ...
in Perm Krai, Russia. It lived about 267 million years ago. The only species is ''Eotitanosuchus olsoni''.


Description

''Eotitanosuchus'' is known from a single large skull without a lower jaw. The skull was , but the overall length may have been over , possibly up to and more than in weight for adult specimens. Like ''
Biarmosuchus ''Biarmosuchus'' is an extinct genus of biarmosuchian therapsids that lived around 267 mya during the Middle Permian period. ''Biarmosuchus'' was discovered in the Perm region of Russia. The first specimen was found in channel sandstone that was ...
tener'', it was primitive in that, though it was a predator, the temple opening behind the eye was small, giving it a weak bite. The temple was, however, larger at the top than in other biarmosuchians.


Paleoecology

''Eotitanosuchus'' fossils were found in the Perm (or Cis-Urals) region of Russia. ''Eotitanosuchus'' was without doubt a dominant animal of its environment. Found preserved in flood deposits (once coastal bogs) containing many skeletons of estemmenosuchids, it has been suggested that this large predator was an excellent swimmer, possibly semi-aquatic or frequenting marshy ground. This, however, is just speculation.


Classification

''Eotitanosuchus'' is often grouped with the
Phthinosuchidae ''Phthinosuchus'' is an extinct genus of therapsids from the Middle Permian of Russia. ''Phthinosuchus'' is the sole member of the family Phthinosuchidae. ''Phthinosuchus'' may have been one of the most primitive therapsids, meaning that its ance ...
and the Biarmosuchidae. In fact, Ivakhnenko (1999) argues that ''
Biarmosuchus ''Biarmosuchus'' is an extinct genus of biarmosuchian therapsids that lived around 267 mya during the Middle Permian period. ''Biarmosuchus'' was discovered in the Perm region of Russia. The first specimen was found in channel sandstone that was ...
tener'' and ''Eotitanosuchus olsoni'' are the same organism, which would eliminate the
Eotitanosuchia Eotitanosuchidae is an extinct family of biarmosuchian therapsids. The Eotitanosuchidae were large predatory therapsids of the Wordian epoch. It was once considered to belong to a separate infraorder of therapsids called Eotitanosuchia. Characte ...
as a separate taxon, though this conclusion does not seem to have been widely accepted. Regardless of the eventual outcome of this debate, Ivakhnenko's paper does seem to show that ''Eotitanosuchus'' is very similar to ''Biarmosuchus''. Further, given the rather close similarity between ''Eotitanosuchus'' and later therapsids, this observation supports the view that Biarmosuchia is
paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...
. Others view ''Eotitanosuchus'' as quite distinct from other basal therapsids and perhaps closer to the Gorgonopsia but gorgonopsian specializations are either not present in ''Eotitanosuchus'' or, as is more often the case, the state of the characters is unknown. This genus is characterized by many primitive features of the septomaxilla, the postorbital, the parietal, the interparietal, the basioccipital, the quadrate rami of the pterygoid and the vomers of the skull. The length of the dorsal process of the premaxilla (front jawbone) and the postorbital twisting (rear side of the skull) constitute specializations that indicate it is not a direct gorgonopsian ancestor. These features, however, are shared by the anteosaur and biarmosuchid lineages.


See also

* List of therapsids


References

* * *
Patricia Vickers-Rich Patricia Arlene Vickers-Rich (born 11 July 1944), also known as Patricia Rich, is an Australian Professor of Palaeontology and Palaeobiology, who researches the environmental changes that have impacted Australia (including the ancient super c ...
and
Thomas H. Rich Thomas may refer to: People * List of people with given name Thomas * Thomas (name) * Thomas (surname) * Saint Thomas (disambiguation) * Thomas Aquinas (1225–1274) Italian Dominican friar, philosopher, and Doctor of the Church * Thomas the Ap ...
, '' The Great Russian Dinosaurs'',
Gunter Graphics Gunter or Günter may refer to: * Gunter rig, a type of rig used in sailing, especially in small boats * Gunter Annex, Alabama, a United States Air Force installation * Gunter, Texas, city in the United States People Surname * Chris Gunter ...
, 1993, pg. 28.


External links


Eotitanosuchidae
at Kheper



at Palaeos * ttp://palaeos.com/vertebrates/therapsida/biarmosuchia.html#Biarmosuchia Therapsida: Biarmosuchiaat Palaeos Biarmosuchians Prehistoric therapsid genera Wordian genera Guadalupian synapsids of Europe Permian Russia Fossils of Russia Fossil taxa described in 1960 {{paleo-therapsid-stub