Emerald Tree Monitor
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The emerald tree monitor (''Varanus prasinus'') or green tree monitor, is a small to medium-sized
arboreal Arboreal locomotion is the locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally, but others are exclusively arboreal. The habitats pose nu ...
monitor lizard. It is known for its unusual coloration, which consists of shades from green to
turquoise Turquoise is an opaque, blue-to-green mineral that is a hydrated phosphate of copper and aluminium, with the chemical formula . It is rare and valuable in finer grades and has been prized as a gemstone and ornamental stone for thousands of year ...
, topped with dark,
transverse Transverse may refer to: *Transverse engine, an engine in which the crankshaft is oriented side-to-side relative to the wheels of the vehicle *Transverse flute, a flute that is held horizontally * Transverse force (or ''Euler force''), the tangen ...
dorsal Dorsal (from Latin ''dorsum'' ‘back’) may refer to: * Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism * Dorsal, positioned on top of an aircraft's fuselage * Dorsal c ...
banding. This coloration helps
camouflage Camouflage is the use of any combination of materials, coloration, or illumination for concealment, either by making animals or objects hard to see, or by disguising them as something else. Examples include the leopard's spotted coat, the b ...
it in its arboreal habitat. Its color also makes the emerald tree monitor highly prized in both the pet trade and zoos alike.


Names

It is known as wbl km in the
Kalam language Kalam is a Kalam language of Papua New Guinea. It is closely related to Kobon, and shares many of the features of that language. Kalam is spoken in Middle Ramu District of Madang Province and in Mount Hagen District of Western Highlands Provin ...
of
Papua New Guinea Papua New Guinea (abbreviated PNG; , ; tpi, Papua Niugini; ho, Papua Niu Gini), officially the Independent State of Papua New Guinea ( tpi, Independen Stet bilong Papua Niugini; ho, Independen Stet bilong Papua Niu Gini), is a country i ...
.


Taxonomy

''Varanus prasinus'' was first described as ''Monitor viridis'' by
John Edward Gray John Edward Gray, FRS (12 February 1800 – 7 March 1875) was a British zoologist. He was the elder brother of zoologist George Robert Gray and son of the pharmacologist and botanist Samuel Frederick Gray (1766–1828). The same is used for ...
in 1831; however, Gray's original
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of sever ...
(RMNH 4812 in the National Natural History Museum in
Leiden Leiden (; in English and archaic Dutch also Leyden) is a city and municipality in the province of South Holland, Netherlands. The municipality of Leiden has a population of 119,713, but the city forms one densely connected agglomeration wi ...
) was lost and the species was redescribed by Schlegel eight years later as ''V. prasinus'' using the found specimen. The
generic Generic or generics may refer to: In business * Generic term, a common name used for a range or class of similar things not protected by trademark * Generic brand, a brand for a product that does not have an associated brand or trademark, other ...
name ''Varanus'' is derived from the
Arabic Arabic (, ' ; , ' or ) is a Semitic language spoken primarily across the Arab world.Semitic languages: an international handbook / edited by Stefan Weninger; in collaboration with Geoffrey Khan, Michael P. Streck, Janet C. E.Watson; Walter ...
word ''waral'' (ورل), which translates to English as "monitor". Its
specific Specific may refer to: * Specificity (disambiguation) * Specific, a cure or therapy for a specific illness Law * Specific deterrence, focussed on an individual * Specific finding, intermediate verdict used by a jury in determining the fina ...
name, ''prasinus'', is
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
for the color
green Green is the color between cyan and yellow on the visible spectrum. It is evoked by light which has a dominant wavelength of roughly 495570 nm. In subtractive color systems, used in painting and color printing, it is created by a combi ...
. ''Varanus prasinus'' is a member of the subgenus ''Euprepiosaurus''. It is closely related to several other
arboreal Arboreal locomotion is the locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally, but others are exclusively arboreal. The habitats pose nu ...
species; when combined, these are often referred to as the ''V. prasinus''
species group In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...
or species complex. In addition to ''V. prasinus'', this species group, whose members are all allopatric, includes '' V. beccarii'' (
Aru Islands The Aru Islands Regency ( id, Kabupaten Kepulauan Aru) is a group of about 95 low-lying islands in the Maluku Islands of eastern Indonesia. It also forms a regency of Maluku Province, with a land area of . At the 2011 Census the Regency had a ...
), '' V. boehmei'' (
Waigeo Island Waigeo is an island in Southwest Papua province of eastern Indonesia. The island is also known as Amberi, or Waigiu. It is the largest of the four main islands in the Raja Ampat Islands archipelago, between Halmahera and about to the north-wes ...
), '' V. bogerti'' ( D'Entrecasteaux Archipelago), '' V. keithhornei'' (
Cape York Peninsula Cape York Peninsula is a large peninsula located in Far North Queensland, Australia. It is the largest unspoiled wilderness in northern Australia.Mittermeier, R.E. et al. (2002). Wilderness: Earth’s last wild places. Mexico City: Agrupació ...
), '' V. kordensis'' ( Biak Island), '' V. macraei'' ( Batanta Island), '' V. reisingeri'' ( Misool Island) and '' V. telenesetes'' (
Rossel Island Rossel Island (named after de Rossel, a senior officer on the French expedition of d'Entrecasteaux, 1791-1793; also known as Yela) is the easternmost island of the Louisiade Archipelago, within the Milne Bay Province of Papua New Guinea. Tree Isl ...
).


Evolutionary history

The family
Varanidae The Varanidae are a family of lizards in the superfamily Varanoidea within the Anguimorpha group. The family, a group of carnivorous and frugivorous lizards, includes the living genus '' Varanus'' and a number of extinct genera more closely rel ...
probably originated in
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an are ...
about 65 million years ago, although some estimates are as early as the late
Mesozoic The Mesozoic Era ( ), also called the Age of Reptiles, the Age of Conifers, and colloquially as the Age of the Dinosaurs is the second-to-last era of Earth's geological history, lasting from about , comprising the Triassic, Jurassic and Cretace ...
(112 million years ago). Monitor lizards probably migrated to Australia and the Indonesian archipelago between 39 and 26 million years ago. All members of the ''V. prasinus'' species group (sometimes referred to as subgenus ''Euprepiosaurus'') are found east of Lydekker’s line in the Indo-Australian Archipelago. This group probably originated in the
Moluccas The Maluku Islands (; Indonesian: ''Kepulauan Maluku'') or the Moluccas () are an archipelago in the east of Indonesia. Tectonically they are located on the Halmahera Plate within the Molucca Sea Collision Zone. Geographically they are located ...
, although dating their origins is complicated by the paucity of the fossil record. The absence of large placental predatory mammals east of
Wallace's line The Wallace Line or Wallace's Line is a faunal boundary line drawn in 1859 by the British naturalist Alfred Russel Wallace and named by English biologist Thomas Henry Huxley that separates the biogeographical realms of Asia and Wallacea, a tr ...
may be partially responsible for the radiation of the ''V. prasinus'' group in the Australo-Papuan region.


Distribution

Emerald tree monitors and their close relatives can be found on the island of
New Guinea New Guinea (; Hiri Motu: ''Niu Gini''; id, Papua, or , historically ) is the world's second-largest island with an area of . Located in Oceania in the southwestern Pacific Ocean, the island is separated from Australia by the wide Torr ...
(split between
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
and
Papua New Guinea Papua New Guinea (abbreviated PNG; , ; tpi, Papua Niugini; ho, Papua Niu Gini), officially the Independent State of Papua New Guinea ( tpi, Independen Stet bilong Papua Niugini; ho, Independen Stet bilong Papua Niu Gini), is a country i ...
) as well as several adjacent islands, and the northern
Torres Strait Islands The Torres Strait Islands are a group of at least 274 small islands in the Torres Strait, a waterway separating far northern continental Australia's Cape York Peninsula and the island of New Guinea. They span an area of , but their total land ...
. The green tree monitor is reported to thrive in lowland environments, including tropical evergreen forests, palm swamps and cocoa plantations.


Description

The emerald tree monitor is about long with a slender body that helps it support itself on narrow branches. It uses its
prehensile Prehensility is the quality of an appendage or organ that has adapted for grasping or holding. The word is derived from the Latin term ''prehendere'', meaning "to grasp". The ability to grasp is likely derived from a number of different orig ...
tail and long claws to grip branches. Unlike other varanids, this monitor defends its tail rather than lashing with it for defense when threatened. The soles of the feet of the emerald tree monitor have enlarged scales which aid the lizard when climbing.


Ecology

When threatened, the emerald tree monitor will flee through vegetation or bite if cornered. It is one of the few social monitors, living in small groups made up of a dominant male, several females, and a few other males and juveniles.


Diet

The emerald tree monitor's diet consists of large tree-dwelling
arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and cuticle made of chiti ...
s, such as katydids, grasshoppers, other
orthopteran Orthoptera () is an order of insects that comprises the grasshoppers, locusts, and crickets, including closely related insects, such as the bush crickets or katydids and wētā. The order is subdivided into two suborders: Caelifera – grassho ...
s, stick insects,
cockroach Cockroaches (or roaches) are a Paraphyly, paraphyletic group of insects belonging to Blattodea, containing all members of the group except termites. About 30 cockroach species out of 4,600 are associated with human habitats. Some species are we ...
es,
beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
s, centipedes,
spider Spiders ( order Araneae) are air-breathing arthropods that have eight legs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species ...
s and crabs, as well as
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
s and small mammals (such as the Moncton's mosaic-tailed rat). Before swallowing stick insects, the lizards tear off the legs.
Captive Captive or Captives may refer to: Arts, entertainment, and media Films * ''Captive'' (1980 film), a sci-fi film, starring Cameron Mitchell and David Ladd * ''Captive'' (1986 film), a British-French film starring Oliver Reed * ''Captive'' (1991 ...
specimens tear off the limbs of rodents prior to eating them; as a result, they are capable of swallowing mammals of a considerable size: A 135-g lizard was documented as eating a 40-g rodent, almost one-third its size. Paleontologist and
Temple University Temple University (Temple or TU) is a public state-related research university in Philadelphia, Pennsylvania. It was founded in 1884 by the Baptist minister Russell Conwell and his congregation Grace Baptist Church of Philadelphia then calle ...
professor Michael Balsai has observed ''V. prasinus'' eating fruit (bananas) in captivity as has herpetologist and author Robert G. Sprackland.


Reproduction

Clutches consist of up to five
eggs Humans and human ancestors have scavenged and eaten animal eggs for millions of years. Humans in Southeast Asia had domesticated chickens and harvested their eggs for food by 1,500 BCE. The most widely consumed eggs are those of fowl, especial ...
, each weighing and measuring about . As many as three clutches are laid throughout the year; clutches have been laid by captives in January, March, April, November, and December. The female emerald tree monitor lays her eggs in arboreal
termite Termites are small insects that live in colonies and have distinct castes (eusocial) and feed on wood or other dead plant matter. Termites comprise the infraorder Isoptera, or alternatively the epifamily Termitoidae, within the order Blatto ...
nests. The eggs hatch between 160 and 190 days later, typically from June to November, after which the young eat the termites and the termite's eggs within minutes of hatching. Sexual maturity is reached in about one year.


References


External links


Photos at Biolib.cz



Mampam.com
{{Taxonbar, from=Q852763 Varanus Monitor lizards of Australia Reptiles of Indonesia Monitor lizards of New Guinea Reptiles of Papua New Guinea Reptiles described in 1839 Taxa named by Hermann Schlegel