Eleotridae
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Eleotridae is a
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
of
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...
commonly known as sleeper gobies, with about 34 genera and 180
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
. Most species are found in the tropical
Indo-Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...
region, but there are also species in
subtropical The subtropical zones or subtropics are geographical zone, geographical and Köppen climate classification, climate zones to the Northern Hemisphere, north and Southern Hemisphere, south of the tropics. Geographically part of the Geographical z ...
and temperate regions, warmer parts of the Americas and near the Atlantic coast in Africa. While many eleotrids pass through a
plankton Plankton are the diverse collection of organisms found in Hydrosphere, water (or atmosphere, air) that are unable to propel themselves against a Ocean current, current (or wind). The individual organisms constituting plankton are called plankt ...
ic stage in the sea and some spend their entire lives in the sea; as adults, the majority live in
freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does include ...
streams and
brackish Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estuari ...
water. One of its genera, '' Caecieleotris'', is
troglobitic A troglobite (or, formally, troglobiont) is an animal species, or population of a species, strictly bound to underground habitats, such as caves. These are separate from species that mainly live in above-ground habitats but are also able to live u ...
. They are especially important as predators in the freshwater stream ecosystems on oceanic islands such as
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...
and
Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only stat ...
that otherwise lack the predatory fish families typical of nearby continents, such as
catfish Catfish (or catfishes; order Siluriformes or Nematognathi) are a diverse group of ray-finned fish. Named for their prominent barbels, which resemble a cat's whiskers, catfish range in size and behavior from the three largest species alive, ...
. Anatomically, they are similar to the gobies (
Gobiidae Gobiidae or gobies is a family of bony fish in the order Gobiiformes, one of the largest fish families comprising more than 2,000 species in more than 200 genera. Most of gobiid fish are relatively small, typically less than in length, and the ...
), though unlike the majority of gobies, they do not have a pelvic sucker.Helfman, G.S., Collette, B.B. & Facey, D.E. (1997): ''The Diversity of Fishes''. Wiley-Blackwell Publishing. p. 264. Like the true gobies, they are generally small fish that live on the substrate, often amongst
vegetation Vegetation is an assemblage of plant species and the ground cover they provide. It is a general term, without specific reference to particular taxa, life forms, structure, spatial extent, or any other specific botanical or geographic character ...
, in burrows, or in crevices within rocks and
coral Corals are marine invertebrates within the class Anthozoa of the phylum Cnidaria. They typically form compact colonies of many identical individual polyps. Coral species include the important reef builders that inhabit tropical oceans and sec ...
reef A reef is a ridge or shoal of rock, coral or similar relatively stable material, lying beneath the surface of a natural body of water. Many reefs result from natural, abiotic processes— deposition of sand, wave erosion planing down rock out ...
s. Although goby-like in many ways, sleeper gobies lack the pelvic fin sucker and that, together with other morphological differences, is used to distinguish the two families. The Gobiidae and Eleotridae likely share a common ancestor and they are both placed in the order
Gobiiformes The Gobiiformes are an order of fish that includes the gobies and their relatives. The order, which was previously considered a suborder of Perciformes, is made up of about 2,211 species that are divided between seven families. Phylogenetic rel ...
, along with a few other small families containing goby-like fishes. '' Dormitator'' and ''
Eleotris ''Eleotris'' is a genus of fish in the family Eleotridae with a circumglobal distribution in tropical and subtropical regions. Species There are currently 27 recognized species in this genus: * '' Eleotris acanthopoma'' Bleeker, 1853 (Spiny ...
'', two of the most widespread and typical genera, include a variety of species that inhabit
marine Marine is an adjective meaning of or pertaining to the sea or ocean. Marine or marines may refer to: Ocean * Maritime (disambiguation) * Marine art * Marine biology * Marine debris * Marine habitats * Marine life * Marine pollution Military * ...
,
estuarine An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environment ...
and freshwater habitats. Among the largest members of the family are predatory species such as the bigmouth sleeper (''Gobiomorus dormitor'') at up to from freshwater near the West Atlantic region and the fat sleeper (''Dormitator maculatus''), which grows to and is widely found in fresh to brackish and shallow marine waters of the southeastern United States and Mexico,Hoedeman, J.J. (1974): ''Naturalists' guide to fresh-water aquarium fish''. Sterling Publishing. p. 1152. However, most are much smaller, such as the fresh- and brackish-water species from Australia and
New Guinea New Guinea (; Hiri Motu Hiri Motu, also known as Police Motu, Pidgin Motu, or just Hiri, is a language of Papua New Guinea, which is spoken in surrounding areas of Port Moresby (Capital of Papua New Guinea). It is a simplified version of ...
, including '' Hypseleotris'', known locally as gudgeons (not to be confused with the Eurasian freshwater cyprinid ''
Gobio gobio ''Gobio gobio'', or the gudgeon, is a species of fish in the family Cyprinidae. This small fish is widely distributed in fresh-water streams and lakes across central and temperate Eurasia. The gudgeon inhabits various fresh-water habitats with ...
'', also known as the gudgeon and after which the Australian sleeper gobies were likely named).Riehl, R. & Baensch, H.A. (1997): ''Aquarium Atlas'' (Volume 2). Voyageur Press. p. 1216. A few of these, such as the empire gudgeon (''H. compressa'') and peacock gudgeon (''Tateurndina ocellicauda''), are sometimes kept in aquariums. The smallest in the family are the Amazonian ''
Leptophilypnion ''Leptophilypnion'' is a genus of tiny fishes in the family Eleotridae endemism, endemic to the Amazon Basin in South America. At less than in standard length they are the smallest sleeper gobies and among the smallest fish. The larger ''Microph ...
'' with a
standard length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish m ...
of less than .


Taxonomy

The family has been divided into three subfamilies: Butinae, Eleotrinae and Milyeringinae. However, because of the deep
divergence In vector calculus, divergence is a vector operator that operates on a vector field, producing a scalar field giving the quantity of the vector field's source at each point. More technically, the divergence represents the volume density of the ...
between the three, some authorities have recommended splitting them into separate families: Butidae, Eleotridae and
Milyeringidae Milyeringidae, the blind cave gobies, is a small family of gobies, in the order Gobiiformes. There are two genera and six species within the family, which is considered to be a subfamily of the Eleotridae by some authorities. Milyeringidae incl ...
.Thacker, C. (2011). Systematics of Butidae and Eleotridae. ''in'' Patzner, R.; J.L. Van Tassell; and M. Kovacic. The Biology of Gobies. Verlag Science Publishers. The 5th edition of
Fishes of the World ''Fishes of the World'' by the American ichthyologist Joseph S. Nelson (1937–2011) is a standard reference for fish systematics. Now in its fifth edition (2016), the work is a comprehensive overview of the diversity and classification of the ...
follows this classification and this means that the following genera are currently included within the Eleotridae. However, the family Xenisthmidae is regarded as a
synonym A synonym is a word, morpheme, or phrase that means exactly or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are all ...
of the Eleotridae, according to the 5th Edition of Fishes of the World. * '' Allomicrodesmus'' Schultz, 1966 * '' Allomogurnda'' Allen, 2003 * '' Belobranchus'' Bleeker, 1857 * '' Bunaka'' Herre, 1927 * '' Caecieleotris'' Walsh & Chakrabarty, 2016 * '' Calumia'' Smith, 1958 * '' Dormitator'' Gill, 1861 * ''
Eleotris ''Eleotris'' is a genus of fish in the family Eleotridae with a circumglobal distribution in tropical and subtropical regions. Species There are currently 27 recognized species in this genus: * '' Eleotris acanthopoma'' Bleeker, 1853 (Spiny ...
'' Bloch & Schneider, 1801 * '' Erotelis'' Poey, 1860 * '' Giuris'' Sauvage, 1880 * ''
Gobiomorphus ''Gobiomorphus'' is a genus of fishes in the family Eleotridae native to New Zealand and Australia. They are typically small, benthic fishes with large, rounded fins and two dorsal fins. Many have an amphidromous lifecycle: the eggs are laid in ...
'' Gill, 1863 * '' Gobiomorus'' Lacépède, 1800 * '' Guavina'' Bleeker, 1874 * '' Gymnoxenisthmus'' Gill, Bogorodsky & Mal, 2014 * '' Hemieleotris'' Meek & Hildebrand, 1916 * '' Hypseleotris'' Gill, 1863 * '' Kimberleyeleotris'' Hoese & Allen, 1987 * ''
Leptophilypnion ''Leptophilypnion'' is a genus of tiny fishes in the family Eleotridae endemism, endemic to the Amazon Basin in South America. At less than in standard length they are the smallest sleeper gobies and among the smallest fish. The larger ''Microph ...
'' Roberts, 2013 * '' Leptophilypnus'' Meek & Hildebrand, 1916 * '' Microphilypnus'' Myers, 1927 * ''
Mogurnda ''Mogurnda'' is a genus of freshwater fishes in the family Eleotridae native to eastern and northern Australia and New Guinea. Several species are endemic to Lake Kutubu in Papua New Guinea. Species The currently recognized species in this genus ...
'' Gill, 1863 * '' Paraxenisthmus'' Gill & Hoese, 1993 * '' Philypnodon'' Bleeker, 1874 * '' Ratsirakia'' Maugé, 1984 * ''
Rotuma Rotuma is a Fijian dependency, consisting of Rotuma Island and nearby islets. The island group is home to a large and unique Polynesian indigenous ethnic group which constitutes a recognisable minority within the population of Fiji, known as " ...
'' Springer, 1988 * '' Tateurndina'' Nichols, 1955 * ''
Tyson Tyson is a male given name of old French origin meaning 'high-spirited', 'fire'. It is from this that a surname arose 'son of Tyson'. Surname *Alan Tyson (1926–2000), British musicologist *Barbara Tyson (born 1964), Canadian actress *Bill Tyso ...
'' Springer, 1983 * '' Xenisthmus'' Snyder, 1908


References

{{Taxonbar, from=Q773959 Gobiiformes Taxa named by Charles Lucien Bonaparte