anatomy Anatomy () is the branch of morphology concerned with the study of the internal structure of organisms and their parts. Anatomy is a branch of natural science that deals with the structural organization of living things. It is an old scien ...

, the extrapyramidal system is a part of the

motor system The motor system is the set of central nervous system, central and peripheral nervous system, peripheral structures in the nervous system that support motor functions, i.e. movement. Peripheral structures may include skeletal muscles and Efferen ...

network Network, networking and networked may refer to: Science and technology * Network theory, the study of graphs as a representation of relations between discrete objects * Network science, an academic field that studies complex networks Mathematics ...

causing involuntary actions. The system is called ''extrapyramidal'' to distinguish it from the tracts of the motor cortex that reach their targets by traveling through the

pyramids A pyramid () is a Nonbuilding structure, structure whose visible surfaces are triangular in broad outline and converge toward the top, making the appearance roughly a Pyramid (geometry), pyramid in the geometric sense. The base of a pyramid ca ...

of the medulla. The

pyramidal tracts The pyramidal tracts include both the corticobulbar tract and the corticospinal tract. These are aggregations of efferent nerve fibers from the upper motor neurons that travel from the cerebral cortex and terminate either in the brainstem (''cort ...

(

corticospinal tract The corticospinal tract is a white matter motor pathway starting at the cerebral cortex that terminates on lower motor neurons and interneurons in the spinal cord, controlling movements of the limbs and trunk. There are more than one million neu ...

and

corticobulbar tract The corticobulbar (or corticonuclear) tract is a two-neuron white matter motor pathway connecting the motor cortex in the cerebral cortex to the Medullary pyramids (brainstem), medullary pyramids, which are part of the brainstem's medulla oblonga ...

s) may directly innervate motor neurons of the spinal cord or brainstem ( anterior (ventral) horn cells or certain cranial nerve nuclei), whereas the extrapyramidal system centers on the modulation and regulation (indirect control) of anterior (ventral) horn cells. __TOC__

Extrapyramidal tracts

The extrapyramidal tracts are chiefly found in the

reticular formation The reticular formation is a set of interconnected nuclei in the brainstem that spans from the lower end of the medulla oblongata to the upper end of the midbrain. The neurons of the reticular formation make up a complex set of neural networks ...

of the

pons The pons (from Latin , "bridge") is part of the brainstem that in humans and other mammals, lies inferior to the midbrain, superior to the medulla oblongata and anterior to the cerebellum. The pons is also called the pons Varolii ("bridge of ...

and medulla, and target

lower motor neuron Lower motor neurons (LMNs) are motor neurons located in either the anterior grey column, anterior nerve roots (spinal lower motor neurons) or the cranial nerve nuclei of the brainstem and cranial nerves with motor function (cranial nerve lower ...

s in the spinal cord that are involved in reflexes, locomotion, complex movements, and postural control. These tracts are in turn modulated by various parts of the central nervous system, including the

nigrostriatal pathway The nigrostriatal pathway is a bilateral dopaminergic pathway in the brain that connects the substantia nigra pars compacta (SNc) in the midbrain with the dorsal striatum (i.e., the caudate nucleus and putamen) in the forebrain. It is one of th ...

, the

basal ganglia The basal ganglia (BG) or basal nuclei are a group of subcortical Nucleus (neuroanatomy), nuclei found in the brains of vertebrates. In humans and other primates, differences exist, primarily in the division of the globus pallidus into externa ...

, the

cerebellum The cerebellum (: cerebella or cerebellums; Latin for 'little brain') is a major feature of the hindbrain of all vertebrates. Although usually smaller than the cerebrum, in some animals such as the mormyrid fishes it may be as large as it or eve ...

, the

vestibular nuclei The vestibular nuclei (VN) are the cranial nuclei for the vestibular nerve located in the brainstem. In Terminologia Anatomica, they are grouped in both the pons and the medulla in the brainstem The brainstem (or brain stem) is the poste ...

, and different sensory areas of the

cerebral cortex The cerebral cortex, also known as the cerebral mantle, is the outer layer of neural tissue of the cerebrum of the brain in humans and other mammals. It is the largest site of Neuron, neural integration in the central nervous system, and plays ...

. All of these regulatory components can be considered part of the extrapyramidal system, in that they modulate motor activity without directly innervating motor neurons. The extrapyramidal tracts include parts of the following: *

rubrospinal tract The rubrospinal tract is one of the descending tracts of the spinal cord. It is a motor control pathway that originates in the red nucleus. It is a part of the lateral indirect extrapyramidal tract. The rubrospinal tract fibers are efferent ne ...

: Conflicts between the motor commands sent by the cerebrum and body position information provided by the proprioceptors cause the cerebellum to stimulate the

red nucleus The red nucleus or nucleus ruber is a structure in the rostral midbrain involved in motor coordination. The red nucleus is pale pink, which is believed to be due to the presence of iron in at least two different forms: hemoglobin and ferritin. ...

of the midbrain. The red nucleus then sends corrective commands to the spinal cord along the rubrospinal tract. * reticulospinal tract: connects the reticular system, a diffuse region of gray matter in the brain stem, to the spinal cord. It also contributes to muscle tone and influences autonomic functions. * lateral vestibulospinal tract: Connects the brain stem nuclei of the vestibular system with the spinal cord. This allows posture, movement, and balance to be modulated on the basis of equilibrium information provided by the vestibular system. * tectospinal tract: This tract projects from the midbrain to the spinal cord and is important for postural movements that are driven by the superior colliculus.

References

{{DEFAULTSORT:Extrapyramidal System Central nervous system Motor system

Extrapyramidal tracts

See also

References