The occipital bone () is a

cranial Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position pro ...

dermal bone and the main bone of the occiput (back and lower part of the

skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, the ...

). It is

trapezoid A quadrilateral with at least one pair of parallel sides is called a trapezoid () in American and Canadian English. In British and other forms of English, it is called a trapezium (). A trapezoid is necessarily a Convex polygon, convex quadri ...

al in shape and curved on itself like a shallow dish. The occipital bone overlies the occipital lobes of the cerebrum. At the base of skull in the occipital bone, there is a large oval opening called the

foramen magnum The foramen magnum ( la, great hole) is a large, oval-shaped opening in the occipital bone of the skull. It is one of the several oval or circular openings (foramina) in the base of the skull. The spinal cord, an extension of the medulla oblon ...

, which allows the passage of the

spinal cord The spinal cord is a long, thin, tubular structure made up of nervous tissue, which extends from the medulla oblongata in the brainstem to the lumbar region of the vertebral column (backbone). The backbone encloses the central canal of the spi ...

. Like the other cranial bones, it is classed as a flat bone. Due to its many attachments and features, the occipital bone is described in terms of separate parts. From its front to the back is the basilar part, also called the basioccipital, at the sides of the foramen magnum are the lateral parts, also called the exoccipitals, and the back is named as the squamous part. The basilar part is a thick, somewhat quadrilateral piece in front of the foramen magnum and directed towards the pharynx. The squamous part is the curved, expanded plate behind the foramen magnum and is the largest part of the occipital bone. Due to its embryonic derivation from paraxial mesoderm (as opposed to neural crest, from which many other craniofacial bones are derived), it has been posited that "the occipital bone as a whole could be considered as a giant vertebra enlarged to support the brain."

Structure

The occipital bone, like the other seven cranial bones, has outer and inner layers (also called ''plates'' or ''tables'') of cortical bone tissue between which is the cancellous bone tissue known in the cranial bones as

diploë Diploë ( or ) is the spongy cancellous bone separating the inner and outer layers of the cortical bone of the skull. In the cranial bones, the layers of compact cortical tissue are familiarly known as the tables of the skull; the outer one is ...

. The bone is especially thick at the ridges, protuberances,

condyles A condyle (;Entry "condyle"
in
basilar part; in the inferior cerebellar fossae it is thin, semitransparent, and without diploë.

Outer surface

Near the middle of the outer surface of the squamous part of the occipital (the largest part) there is a prominence – the external occipital protuberance. The highest point of this is called the inion. From the inion, along the midline of the squamous part until the foramen magnum, runs a ridge – the external occipital crest (also called the medial nuchal line) and this gives attachment to the nuchal ligament. Running across the outside of the occipital bone are three curved lines and one line (the medial line) that runs down to the foramen magnum. These are known as the nuchal lines which give attachment to various ligaments and muscles. They are named as the ''highest'', ''superior'' and ''inferior'' nuchal lines. The inferior nuchal line runs across the midpoint of the ''median'' nuchal line. The area above the highest nuchal line is termed the occipital plane and the area below this line is termed the nuchal plane.

Inner surface

The inner surface of the occipital bone forms the base of the posterior cranial fossa. The

is a large hole situated in the middle, with the clivus, a smooth part of the occipital bone travelling upwards in front of it. The median internal occipital crest travels behind it to the internal occipital protuberance, and serves as a point of attachment to the falx cerebri. To the sides of the foramen sitting at the junction between the lateral and base of the occipital bone are the hypoglossal canals. Further out, at each junction between the occipital and petrous portion of the temporal bone lies a jugular foramen. The inner surface of the occipital bone is marked by dividing lines as shallow ridges, that form four fossae or depressions. The lines are called the cruciform (cross-shaped)

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. At the midpoint where the lines intersect a raised part is formed called the internal occipital protuberance. From each side of this eminence runs a groove for the transverse sinuses. There are two midline skull landmarks at the

. The basion is the most anterior point of the opening and the opisthion is the point on the opposite posterior part. The basion lines up with the dens.

Foramen magnum

The foramen magnum ( la, large hole) is a large oval foramen longest front to back; it is wider behind than in front where it is encroached upon by the occipital condyles. The clivus, a smooth bony section, travels upwards on the front surface of the foramen, and the median internal occipital crest travels behind it. Through the foramen passes the

medulla oblongata The medulla oblongata or simply medulla is a long stem-like structure which makes up the lower part of the brainstem. It is anterior and partially inferior to the cerebellum. It is a cone-shaped neuronal mass responsible for autonomic (involun ...

and its membranes, the accessory nerves, the vertebral arteries, the

anterior Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position prov ...

and

posterior spinal arteries The posterior spinal artery (dorsal spinal arteries) arises from the vertebral artery in 25% of humans or the posterior inferior cerebellar artery in 75% of humans, adjacent to the medulla oblongata. It supplies the grey and white posterior columns ...

, and the tectorial membrane and alar ligaments.

Angles

The superior angle of the occipital bone articulates with the occipital angles of the parietal bones and, in the fetal skull, corresponds in position with the posterior fontanelle. The lateral angles are situated at the extremities of the groove for the transverse sinuses: each is received into the interval between the mastoid angle of the parietal bone, and the mastoid portion of the temporal bone. The inferior angle is fused with the body of the sphenoid bone.

Borders

The superior borders extend from the superior to the lateral angles: they are deeply serrated for articulation with the occipital borders of the parietals, and form by this union the lambdoidal suture. The inferior borders extend from the lateral angles to the inferior angle; the upper half of each articulates with the mastoid portion of the corresponding temporal, the lower half with the petrous part of the same bone. These two portions of the inferior border are separated from one another by the jugular process, the notch on the anterior surface of which forms the posterior part of the jugular foramen.

Sutures

File:Lambdoid suture.png, Lambdoid suture File:Occipitomastoid suture.png, Occipitomastoid suture The lambdoid suture joins the occipital bone to the parietal bones. The occipitomastoid suture joins the occipital bone and mastoid portion of the temporal bone. The sphenobasilar suture joins the basilar part of the occipital bone and the back of the

sphenoid bone The sphenoid bone is an unpaired bone of the neurocranium. It is situated in the middle of the skull towards the front, in front of the basilar part of occipital bone, basilar part of the occipital bone. The sphenoid bone is one of the seven bones ...

body . The petrous-basilar suture joins the side edge of the basilar part of the occipital bone to the petrous-part of the temporal bone .

Development

The occipital plane

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of the squamous part of the occipital bone is developed in membrane, and may remain separate throughout life when it constitutes the interparietal bone; the rest of the bone is developed in cartilage. The number of nuclei for the occipital plane is usually given as four, two appearing near the middle line about the second month, and two some little distance from the middle line about the third month of fetal life. The nuchal plane of the squamous part is ossified from two centers, which appear about the seventh week of fetal life and soon unite to form a single piece. Union of the upper and lower portions of the squamous part takes place in the third month of fetal life. An occasional centre ( Kerckring) appears in the posterior margin of the foramen magnum during the fifth month; this forms a separate ossicle (sometimes double) which unites with the rest of the squamous part before birth. Each of the lateral parts begins to ossify from a single center during the eighth week of fetal life. The basilar portion is ossified from two centers, one in front of the other; these appear about the sixth week of fetal life and rapidly coalesce. The occipital plane is said to be ossified from two centers and the basilar portion from one. About the fourth year the squamous part and the two lateral parts unite, and by about the sixth year the bone consists of a single piece. Between the 18th and 25th years the occipital and

become united, forming a single bone.

Clinical significance

Trauma to the occiput can cause a fracture of the base of the skull, called a basilar skull fracture. The ''basion-dens line'' as seen on a radiograph is the distance between the basion and the top of the dens, used in the diagnosis of dissociation injuries. Genetic disorders can cause a prominent occiput as found in Edwards syndrome, and

Beckwith–Wiedemann syndrome Beckwith–Wiedemann syndrome (; abbreviated BWS) is an overgrowth disorder usually present at birth, characterized by an increased risk of childhood cancer and certain congenital features. A minority (97th centile) * Macroglossia * Hemihyperplas ...

. The identification of the location of the fetal occiput is important in delivery.

Etymology

Occipital stems from Latin occiput "back of the skull", from ob "against, behind" + caput "head". Distinguished from

sinciput In human anatomy, the forehead is an area of the head bounded by three features, two of the skull and one of the scalp. The top of the forehead is marked by the hairline, the edge of the area where hair on the scalp grows. The bottom of the fore ...

(anterior part of the skull)."occipital" A Dictionary of Zoology. Ed. Michael Allaby. Oxford University Press 2009

Other animals

In many animals these parts stay separate throughout life; for example, in the dog as four parts: squamous part (supraoccipital); lateral parts–left and right parts (exoccipital); basilar part (basioccipital). The occipital bone is part of the

endocranium The endocranium in comparative anatomy is a part of the skull base in vertebrates and it represents the basal, inner part of the cranium. The term is also applied to the outer layer of the dura mater in human anatomy. Structure Structurally, t ...

, the most basal portion of the skull. In Chondrichthyes and Agnatha, the occipital does not form as a separate element, but remains part of the

chondrocranium The chondrocranium (or ''cartilaginous neurocranium'') is the primitive cartilage, cartilaginous skeleton, skeletal structure of the fetal skull that grows to envelop the rapidly growing embryonic brain.Salentijn, L. ''Biology of Mineralized Tissue ...

throughout life. In most higher vertebrates, the foramen magnum is surrounded by a ring of four bones. The basioccipital lies in front of the opening, the two exoccipital condyles lie to either side, and the larger supraoccipital lies to the posterior, and forms at least part of the rear of the cranium. In many

bony fish Osteichthyes (), popularly referred to as the bony fish, is a diverse superclass of fish that have skeletons primarily composed of bone tissue. They can be contrasted with the Chondrichthyes, which have skeletons primarily composed of cartilag ...

and

amphibian Amphibians are tetrapod, four-limbed and ectothermic vertebrates of the Class (biology), class Amphibia. All living amphibians belong to the group Lissamphibia. They inhabit a wide variety of habitats, with most species living within terres ...

s, the supraoccipital is never ossified, and remains as

cartilage Cartilage is a resilient and smooth type of connective tissue. In tetrapods, it covers and protects the ends of long bones at the joints as articular cartilage, and is a structural component of many body parts including the rib cage, the neck an ...

throughout life. In primitive forms the basioccipital and exoccipitals somewhat resemble the centrum and neural arches of a vertebra, and form in a similar manner in the embryo. Together, these latter bones usually form a single concave circular condyle for the articulation of the first vertebra. In

mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...

s, however, the condyle has divided in two, a pattern otherwise seen only in a few amphibians. Most mammals also have a single fused occipital bone, formed from the four separate elements around the foramen magnum, along with the paired postparietal bones that form the rear of the cranial roof in other vertebrates.

Additional images

File:Occipital bone animation.gif, Position of occipital bone (shown in green). Animation. File:Occipital bone animation2.gif, Outer surface File:Occipital bone animation3.gif, Inner surface. Frontal bone and parietal bones are removed. File:Occipital bone.jpg, Occipital bone File:Occipital bone 2.jpg, Occipital bone File:Gray308.png, Median sagittal section through the occipital bone and first three cervical vertebræ File:Basilar part.jpg, Basilar part File:Slide4llll.JPG, Occipital bone