In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, which completely lose their foliage during the winter or dry season.

Evergreen species

There are many different kinds of evergreen plants, both trees and

shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...

s. Evergreens include: *Most species of

conifers Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All extan ...

(e.g., pine, hemlock, blue spruce, and red cedar), but not all (e.g., larch) * Live oak, holly, and "ancient" gymnosperms such as cycads *Most

angiosperm Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...

s from frost-free climates, and

rainforest Rainforests are characterized by a closed and continuous tree canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforest can be classified as tropical rainforest or temperate rainfores ...

trees *All Eucalypts * Clubmosses and relatives * Bamboos The Latin binomial term , meaning "always green", refers to the evergreen nature of the plant, for instance :''

Cupressus sempervirens ''Cupressus sempervirens'', the Mediterranean cypress (also known as Italian cypress, Tuscan cypress, Persian cypress, or pencil pine), is a species of cypress native to the eastern Mediterranean region, in northeast Libya, southern Albania, sou ...

'' (a cypress) :'' Lonicera sempervirens'' (a honeysuckle) :'' Sequoia sempervirens'' (a sequoia) Leaf longevity in evergreen plants varies from a few months to several decades (over thirty years in the

Great Basin bristlecone pine ''Pinus longaeva'' (commonly referred to as the Great Basin bristlecone pine, intermountain bristlecone pine, or western bristlecone pine) is a long-living species of bristlecone pine tree found in the higher mountains of California, Nevada, and ...

Evergreen families

Japanese umbrella pine is unique in that it has its own family of which it is the only species.

Differences between evergreen and deciduous species

Evergreen and deciduous species vary in a range of morphological and physiological characters. Generally, broad-leaved evergreen species have thicker leaves than deciduous species, with a larger volume of parenchyma and air spaces per unit leaf area. They have larger leaf biomass per unit leaf area, and hence a lower

specific leaf area Specific leaf area (SLA) is the ratio of leaf area to leaf dry mass. The inverse of SLA is Leaf Mass per Area (LMA). Rationale Specific leaf area is a ratio indicating how much leaf area a plant builds with a given amount of leaf biomass: SLA \ = ...

. Construction costs do not differ between the groups. Evergreens have generally a larger fraction of total plant biomass present as leaves (LMF), but they often have a lower rate of photosynthesis.

Reasons for being evergreen or deciduous

Deciduous trees shed their leaves usually as an adaptation to a cold or dry/wet season. Evergreen trees also lose leaves, but each tree loses its leaves gradually and not all at once. Most tropical rainforest plants are considered to be evergreens, replacing their leaves gradually throughout the year as the leaves age and fall, whereas species growing in seasonally arid climates may be either evergreen or deciduous. Most warm temperate climate plants are also evergreen. In cool temperate climates, fewer plants are evergreen. In this climate, there is a predominance of

because few evergreen broadleaf plants can tolerate severe cold below about . In areas where there is a reason for being deciduous, e.g. a cold season or dry season, evergreen plants are usually an adaptation of low nutrient levels. Additionally, they usually have hard leaves and have an excellent water economy due to scarce resources in the area in which they reside. The excellent water economy within the evergreen species is due to high abundance when compared to deciduous species. Whereas deciduous trees lose nutrients whenever they lose their leaves. In warmer areas, species such as some pines and

cypresses Cypress is a common name for various coniferous trees or shrubs of northern temperate regions that belong to the family Cupressaceae. The word ''cypress'' is derived from Old French ''cipres'', which was imported from Latin ''cypressus'', the la ...

grow on poor soils and disturbed ground. In ''

Rhododendron ''Rhododendron'' (; from Ancient Greek ''rhódon'' "rose" and ''déndron'' "tree") is a very large genus of about 1,024 species of woody plants in the heath family (Ericaceae). They can be either evergreen or deciduous. Most species are nati ...

'', a genus with many broadleaf evergreens, several species grow in mature forests but are usually found on highly acidic soil where the nutrients are less available to plants. In taiga or boreal forests, it is too cold for the

organic matter in the soil Soil organic matter (SOM) is the organic matter component of soil, consisting of plant and animal detritus at various stages of decomposition, cells and tissues of soil microbes, and substances that soil microbes synthesize. SOM provides numerous ...

to decay rapidly, so the nutrients in the soil are less easily available to plants, thus favoring evergreens. In temperate climates, evergreens can reinforce their own survival; evergreen leaf and needle litter has a higher carbon-nitrogen ratio than deciduous leaf litter, contributing to a higher soil acidity and lower soil nitrogen content. This is the case with Mediterranean evergreen seedlings, which have unique C and N storages that allow stored resources to determine fast growth within the species, limiting competition and bolstering survival. These conditions favor the growth of more evergreens and make it more difficult for deciduous plants to persist. In addition, the shelter provided by existing evergreen plants can make it easier for younger evergreen plants to survive cold and/or drought.Sobrado, M. A. (1991) "Cost-Benefit Relationships in Deciduous and Evergreen Leaves of Tropical Dry Forest Species". ''Functional Ecology'' 5 (5): 608–616.

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