Eurypygae
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Eurypygiformes is an order formed by the kagus, comprising two species in the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Rhynochetidae ''Rhynochetos'' is a genus of ground-dwelling birds in the monotypic family Rhynochetidae. It contains two species, both endemic to New Caledonia, one of which is extinct. Taxonomy ''Rhynochetos jubatus'' is the only extant species of this gen ...
endemic to
New Caledonia ) , anthem = "" , image_map = New Caledonia on the globe (small islands magnified) (Polynesia centered).svg , map_alt = Location of New Caledonia , map_caption = Location of New Caledonia , mapsize = 290px , subdivision_type = Sovereign st ...
, and the
sunbittern The sunbittern (''Eurypyga helias'') is a bittern-like bird of tropical regions of the Americas, and the sole member of the family Eurypygidae (sometimes spelled Eurypigidae) and genus ''Eurypyga''. It is found in Central and South America, and ...
(''Eurypyga helias'') from the tropical regions of the
Americas The Americas, which are sometimes collectively called America, are a landmass comprising the totality of North and South America. The Americas make up most of the land in Earth's Western Hemisphere and comprise the New World. Along with th ...
. Its closest relatives appear to be the tropicbirds of the tropical Atlantic, Indian, and Pacific oceans.


Classification

The affinities of Eurypygiformes are not very well resolved. The group consists of two families from a
Gondwana Gondwana () was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic (about 550 million years ago) and began to break up during the Jurassic period (about 180 million years ago). The final stages ...
n lineage of birds. Based on some morphological characteristics, they were initially classed as members of the family
Ardeidae The herons are long-legged, long-necked, freshwater and coastal birds in the family Ardeidae, with 72 recognised species, some of which are referred to as egrets or bitterns rather than herons. Members of the genera ''Botaurus'' and ''Ixobrychus ...
, and later the
Gruiformes The Gruiformes are an order (biology), order containing a considerable number of living and extinct bird family (biology), families, with a widespread geographical diversity. Gruiform means "crane-like". Traditionally, a number of wading and t ...
. According to Jarvis, et al.'s 2014 "Whole-genome analyses resolve early branches in the tree of life of modern birds", the group is distantly related to the
Phaethontiformes The Phaethontiformes are an order of birds. They contain one extant family, the tropicbirds (Phaethontidae), and one extinct family Prophaethontidae from the early Cenozoic. Several fossil genera have been described. The tropicbirds were tradit ...
.Whole-genome analyses resolve early branches in the tree of life of modern birds
Erich D. Jarvis, ''et al''., ''Science'' 12 December 2014: Vol. 346 no. 6215 pp. 1320-1331 DOI: 10.1126/science.1253451
When seen as a gruiform, the
kagu The kagu or cagou (''Rhynochetos jubatus'') is a crested, long-legged, and bluish-grey bird endemic to the dense mountain forests of New Caledonia. It is the only surviving member of the genus ''Rhynochetos'' and the family Rhynochetidae, alt ...
is generally considered related to the extinct
adzebill The adzebills, genus ''Aptornis'', were two closely related bird species, the North Island adzebill, (''Aptornis otidiformis''), and the South Island adzebill, (''Aptornis defossor''), of the extinct family Aptornithidae. The family was endemic ...
s from
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...
and the
sunbittern The sunbittern (''Eurypyga helias'') is a bittern-like bird of tropical regions of the Americas, and the sole member of the family Eurypygidae (sometimes spelled Eurypigidae) and genus ''Eurypyga''. It is found in Central and South America, and ...
from
Central Central is an adjective usually referring to being in the center of some place or (mathematical) object. Central may also refer to: Directions and generalised locations * Central Africa, a region in the centre of Africa continent, also known as ...
and
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
. Recent studies do indicate that the sunbittern is the closest living relative of the
kagu The kagu or cagou (''Rhynochetos jubatus'') is a crested, long-legged, and bluish-grey bird endemic to the dense mountain forests of New Caledonia. It is the only surviving member of the genus ''Rhynochetos'' and the family Rhynochetidae, alt ...
. For example, Fain & Houde found these to be certainly
sister taxa In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and t ...
and Furo ''et al.'' indicated a close phylogenetic relationship between them by cytotaxonomy; they suggest that their common ancestor was separated by the Gondwana vicariance in South America and New Caledonia, respectively. They and the mesites did not group with traditional Gruiformes in their study, but instead with their proposed
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
Metaves Metaves ("other birds") is a controversial group proposed by Fain & Houde (2004) and later rescued on the studies of Ericson ''et al.'' (2006)Ericson, P.G.P. ''et al.'' (2006Diversification of Neoaves: integration of molecular sequence data and f ...
, which also includes the
hoatzin The hoatzin ( ) or hoactzin ( ), (''Opisthocomus hoazin''), is the only species in the order Opisthocomiformes. It is a species of tropical bird found in swamps, riparian forests, and mangroves of the Amazon and the Orinoco basins in South Ameri ...
,
pigeons Columbidae () is a bird family consisting of doves and pigeons. It is the only family in the order Columbiformes. These are stout-bodied birds with short necks and short slender bills that in some species feature fleshy ceres. They primarily ...
,
Caprimulgiformes Nightjars are medium-sized nocturnal or crepuscular birds in the family Caprimulgidae and order Caprimulgiformes, characterised by long wings, short legs, and very short bills. They are sometimes called goatsuckers, due to the ancient folk tal ...
,
flamingos Flamingos or flamingoes are a type of wading bird in the family Phoenicopteridae, which is the only extant family in the order Phoenicopteriformes. There are four flamingo species distributed throughout the Americas (including the Caribbean) ...
,
tropicbirds Tropicbirds are a family, Phaethontidae, of tropical pelagic seabirds. They are the sole living representatives of the order Phaethontiformes. For many years they were considered part of the Pelecaniformes, but genetics indicates they are most cl ...
,
Apodiformes Traditionally, the bird order Apodiformes contained three living families: the swifts (Apodidae), the treeswifts (Hemiprocnidae), and the hummingbirds (Trochilidae). In the Sibley-Ahlquist taxonomy, this order is raised to a superorder Apodi ...
,
sandgrouse Sandgrouse is the common name for Pteroclidae , a family (biology), family of sixteen species of bird, members of the order Pterocliformes . They are traditionally placed in two Genus, genera. The two central Asian species are classified as ''Sy ...
and
grebes Grebes () are aquatic diving birds in the order (biology), order Podicipediformes . Grebes are widely distributed freshwater birds, with some species also found in sea, marine habitats during Bird migration, migration and winter. Some flightless ...
. The internal structure of this group was not well resolvable by their data, and contains numerous groupings not otherwise corroborated (such as Caprimulgidae and
flamingos Flamingos or flamingoes are a type of wading bird in the family Phoenicopteridae, which is the only extant family in the order Phoenicopteriformes. There are four flamingo species distributed throughout the Americas (including the Caribbean) ...
). The usefulness and monophyly of "Metaves" is therefore unclear. Notwithstanding, the
kagu The kagu or cagou (''Rhynochetos jubatus'') is a crested, long-legged, and bluish-grey bird endemic to the dense mountain forests of New Caledonia. It is the only surviving member of the genus ''Rhynochetos'' and the family Rhynochetidae, alt ...
and sunbittern – and possibly the adzebills – seem to form a distinct
Gondwana Gondwana () was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic (about 550 million years ago) and began to break up during the Jurassic period (about 180 million years ago). The final stages ...
n lineage of birds, possibly one order, possibly more, even though the relationships between them, the mesites, and the "core Gruiformes" are not yet resolved. It is notable, however, that the sunbittern and the mesites possess
powder down The down of birds is a layer of fine feathers found under the tougher exterior feathers. Very young birds are clad only in down. Powder down is a specialized type of down found only in a few groups of birds. Down is a fine thermal insulator and ...
, whereas the "core Gruiformes" do not. While the
kagu The kagu or cagou (''Rhynochetos jubatus'') is a crested, long-legged, and bluish-grey bird endemic to the dense mountain forests of New Caledonia. It is the only surviving member of the genus ''Rhynochetos'' and the family Rhynochetidae, alt ...
is the only living species in the family Rhynochetidae, a larger species, the
lowland kagu The lowland kagu (''Rhynochetos orarius'') is a large, extinct species of kagu. It was endemic to the island of New Caledonia in Melanesia in the south-west Pacific region. It was described from subfossil bones found at the Pindai Caves paleon ...
(''Rhynochetos orarius''), has been described from late
Holocene The Holocene ( ) is the current geological epoch. It began approximately 11,650 cal years Before Present (), after the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene togethe ...
subfossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in ...
remains. The body measurements of this species were 15% greater than '' Rhynochetos jubatus'', with no overlap in measurements except those of the wings. Given that the sites where ''R. orarius'' has been found are all lowland sites, and that no fossils of ''R. jubatus'' were found in these sites, the scientists who described the fossils suggested they represented highland and lowland species, respectively. ''R. orarius'' is one of many species to have become extinct in New Caledonia after the arrival of humans. The validity of the species has been questioned by some authors,del Hoyo, J. Elliott, A. & Sargatal, J. (editors). (1996) ''Handbook of the Birds of the World. Volume 3: Hoatzin to Auks''. Lynx Edicions. but accepted by others. Fossil
Messelornithidae Messelornithidae is an extinct clade of gruiform birds, closely related to modern rails. The fossil record are from the Paleocene to the early Oligocene of Europe and North America North America is a continent in the Northern Hemisphe ...
once included in Eurypygiformes are now considered the oldest known members of Ralloidea (
Gruiformes The Gruiformes are an order (biology), order containing a considerable number of living and extinct bird family (biology), families, with a widespread geographical diversity. Gruiform means "crane-like". Traditionally, a number of wading and t ...
).


References

{{Taxonbar, from=Q147907 Neognathae Bird orders