European Pine Marten
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The European pine marten (''Martes martes''), also known as the pine marten, is a
mustelid The Mustelidae (; from Latin , weasel) are a diverse family of carnivoran mammals, including weasels, badgers, otters, polecats, martens, grisons, and wolverines. Otherwise known as mustelids (), they form the largest family in the suborde ...
native to and widespread in most of
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
,
Asia Minor Anatolia (), also known as Asia Minor, is a peninsula in West Asia that makes up the majority of the land area of Turkey. It is the westernmost protrusion of Asia and is geographically bounded by the Mediterranean Sea to the south, the Aegean ...
, the
Caucasus The Caucasus () or Caucasia (), is a region spanning Eastern Europe and Western Asia. It is situated between the Black Sea and the Caspian Sea, comprising parts of Southern Russia, Georgia, Armenia, and Azerbaijan. The Caucasus Mountains, i ...
, and parts of
Iran Iran, officially the Islamic Republic of Iran (IRI) and also known as Persia, is a country in West Asia. It borders Iraq to the west, Turkey, Azerbaijan, and Armenia to the northwest, the Caspian Sea to the north, Turkmenistan to the nort ...
,
Iraq Iraq, officially the Republic of Iraq, is a country in West Asia. It is bordered by Saudi Arabia to Iraq–Saudi Arabia border, the south, Turkey to Iraq–Turkey border, the north, Iran to Iran–Iraq border, the east, the Persian Gulf and ...
, and
Syria Syria, officially the Syrian Arab Republic, is a country in West Asia located in the Eastern Mediterranean and the Levant. It borders the Mediterranean Sea to the west, Turkey to Syria–Turkey border, the north, Iraq to Iraq–Syria border, t ...
. It is listed as
Least Concern A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wil ...
on the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is an inventory of the global conservation status and extinction risk of biological ...
. It is less commonly known as baum marten or sweet marten.


Description

The European pine marten's fur is usually light to dark brown. It is short and coarse in the summer, growing longer and silkier during the winter. It has a cream- to yellow-coloured "bib" marking on its throat. Its body is up to long, with a bushy tail of about . It weighs around ; males are slightly larger than females. It has excellent senses of sight, smell, and hearing.


Distribution and habitat


Britain and Ireland

The European pine marten was for many years common only in northwestern
Scotland Scotland is a Countries of the United Kingdom, country that is part of the United Kingdom. It contains nearly one-third of the United Kingdom's land area, consisting of the northern part of the island of Great Britain and more than 790 adjac ...
. A study in 2012 found that it has spread from the
Scottish Highlands The Highlands (; , ) is a historical region of Scotland. Culturally, the Highlands and the Scottish Lowlands, Lowlands diverged from the Late Middle Ages into the modern period, when Scots language, Lowland Scots language replaced Scottish Gae ...
north into
Sutherland Sutherland () is a Counties of Scotland, historic county, registration county and lieutenancy areas of Scotland, lieutenancy area in the Scottish Highlands, Highlands of Scotland. The name dates from the Scandinavian Scotland, Viking era when t ...
and
Caithness Caithness (; ; ) is a Shires of Scotland, historic county, registration county and Lieutenancy areas of Scotland, lieutenancy area of Scotland. There are two towns, being Wick, Caithness, Wick, which was the county town, and Thurso. The count ...
and southeastwards from the Great Glen into
Moray Moray ( ; or ) is one of the 32 council areas of Scotland. It lies in the north-east of the country, with a coastline on the Moray Firth, and borders the council areas of Aberdeenshire and Highland. Its council is based in Elgin, the area' ...
,
Aberdeenshire Aberdeenshire (; ) is one of the 32 Subdivisions of Scotland#council areas of Scotland, council areas of Scotland. It takes its name from the Shires of Scotland, historic county of Aberdeenshire (historic), Aberdeenshire, which had substantial ...
,
Perthshire Perthshire (Scottish English, locally: ; ), officially the County of Perth, is a Shires of Scotland, historic county and registration county in central Scotland. Geographically it extends from Strathmore, Angus and Perth & Kinross, Strathmore ...
, Tayside, and
Stirlingshire Stirlingshire or the County of Stirling ( ) is a Shires of Scotland, historic county and registration county of Scotland. Its county town is Stirling.Registers of Scotland. Publications, leaflets, Land Register Counties. It borders Perthshir ...
, with some in the
Central Belt The Central Belt of Scotland is the Demographics of Scotland, area of highest population density within Scotland. Depending on the definition used, it has a population of between 2.4 and 4.2 million (the country's total was around 5.4 million in ...
, on the
Kintyre Kintyre (, ) is a peninsula in western Scotland, in the southwest of Argyll and Bute. The peninsula stretches about , from the Mull of Kintyre in the south to East Loch Tarbert, Argyll, East and West Loch Tarbert, Argyll, West Loch Tarbert in t ...
and Cowal peninsulas, and on the islands of Skye and Mull. The expansion in the Galloway Forest has been limited compared with that in the core marten range. Martens were reintroduced to the Glen Trool Forest in the early 1980s and only restricted spread has occurred from there. In
England England is a Countries of the United Kingdom, country that is part of the United Kingdom. It is located on the island of Great Britain, of which it covers about 62%, and List of islands of England, more than 100 smaller adjacent islands. It ...
, the European pine marten was considered to be extinct. Analysis of a scat found at Kidland Forest in
Northumberland Northumberland ( ) is a ceremonial counties of England, ceremonial county in North East England, on the Anglo-Scottish border, border with Scotland. It is bordered by the North Sea to the east, Tyne and Wear and County Durham to the south, Cumb ...
in June 2010 may represent either a recolonisation from Scotland, or a relict population that has escaped notice previously. There have been numerous reported sightings of European pine martens in
Cumbria Cumbria ( ) is a ceremonial county in North West England. It borders the Scottish council areas of Dumfries and Galloway and Scottish Borders to the north, Northumberland and County Durham to the east, North Yorkshire to the south-east, Lancash ...
over the years; however, only in 2011, was scat was found and DNA-tested, followed in October 2022 by images of a European pine marten in Grizedale Forest. In July 2015, the first confirmed sighting of a European pine marten in England for over a century was recorded by an amateur photographer in the woodlands of
Shropshire Shropshire (; abbreviated SalopAlso used officially as the name of the county from 1974–1980. The demonym for inhabitants of the county "Salopian" derives from this name.) is a Ceremonial counties of England, ceremonial county in the West M ...
. Sightings have continued in the area and juveniles were recorded in 2019, indicating a breeding population. In July 2017, footage of a live European pine marten was captured by a camera trap in the
North York Moors The North York Moors is an upland area in north-eastern Yorkshire, England. It contains one of the largest expanses of Calluna, heather moorland in the United Kingdom. The area was designated as a national parks of England and Wales, National P ...
in
Yorkshire Yorkshire ( ) is an area of Northern England which was History of Yorkshire, historically a county. Despite no longer being used for administration, Yorkshire retains a strong regional identity. The county was named after its county town, the ...
; in March 2018, the first ever footage of a European pine marten in Northumberland was captured by the Back from the Brink pine marten project. The European pine marten's presence in Southern England's
New Forest The New Forest is one of the largest remaining tracts of unenclosed pasture land, heathland and forest in Southern England, covering southwest Hampshire and southeast Wiltshire. It was proclaimed a royal forest by William the Conqueror, featu ...
was confirmed in 2021, and they are now breeding there. In September 2022, the first European pine marten to be seen in London in a century was spotted by the
Zoological Society of London The Zoological Society of London (ZSL) is a charity and organization devoted to the worldwide animal conservation, conservation of animals and their habitat conservation, habitats. It was founded in 1826. Since 1828, it has maintained London Zo ...
's wildlife cameras as part of a hedgehog monitoring program. The Kent Wildlife Trust in collaboration with the
Sussex Wildlife Trust The Sussex Wildlife Trust (SWT) is a conservation charity which aims to protect natural life in Sussex. It was founded in 1961 and is one of 46 The Wildlife Trusts partnership, wildlife trusts across the UK and the Isle of Man and Alderney. As ...
announced a project in February 2024 to assess the feasibility of reintroducing European pine martens to
South East England South East England is one of the nine official regions of England, regions of England that are in the ITL 1 statistical regions of England, top level category for Statistics, statistical purposes. It consists of the nine counties of england, ...
. There is a small population of the European pine marten in
Wales Wales ( ) is a Countries of the United Kingdom, country that is part of the United Kingdom. It is bordered by the Irish Sea to the north and west, England to the England–Wales border, east, the Bristol Channel to the south, and the Celtic ...
. Scat found in Cwm Rheidol forest in 2007 was confirmed by DNA testing to be from a European pine marten. A male was found in 2012 as road kill near
Newtown, Powys Newtown () is a town in Powys, Wales. It lies on the River Severn in the community of Newtown and Llanllwchaiarn, within the Historic counties of Wales, historic boundaries of Montgomeryshire. It was designated a New towns movement, new town in ...
. This was the first confirmation in Wales of the species, living or dead, since 1971. The Vincent Wildlife Trust (VWT) has begun a reinforcement of these mammals in the mid-Wales area. During autumn 2015, 20 European pine martens were captured in Scotland, in areas where a healthy European pine marten population occurs, under licence from Scottish Natural Heritage. These individuals were translocated and released in an area of mid-Wales. All of the martens were fitted with radio collars and are being tracked daily to monitor their movements and find out where they have set up territories. During autumn 2016, the VWT planned to capture and release another 20 European pine martens in the hope of creating a self-sustaining population. Following the success of these projects, 35 European pine martens were reintroduced between 2019 and 2021 to the
Forest of Dean The Forest of Dean is a geographical, historical and cultural region in the western part of the Counties of England, county of Gloucestershire, England. It forms a roughly triangle, triangular plateau bounded by the River Wye to the west and no ...
in
South West England South West England, or the South West of England, is one of the nine official regions of England, regions of England in the United Kingdom. Additionally, it is one of four regions that altogether make up Southern England. South West England con ...
where they were last seen in 1860, with the hope that in time they will merge with the growing population in Wales to form a contiguous presence. After successful breeding every year, by 2023 their numbers were estimated to be approaching 60. The European pine marten is still quite rare in
Ireland Ireland (, ; ; Ulster Scots dialect, Ulster-Scots: ) is an island in the North Atlantic Ocean, in Northwestern Europe. Geopolitically, the island is divided between the Republic of Ireland (officially Names of the Irish state, named Irelan ...
, but the population is recovering and spreading; its traditional strongholds are in the west and south, especially
the Burren The Burren ( ; ) is a karst/glaciokarst landscape centred in County Clare, on the west coast of Republic of Ireland, Ireland.
and Killarney National Park, but the population in the Midlands has significantly increased in recent years. A study managed by academics at
Queen's University Belfast The Queen's University of Belfast, commonly known as Queen's University Belfast (; abbreviated Queen's or QUB), is a public research university in Belfast, Northern Ireland, United Kingdom. The university received its charter in 1845 as part of ...
, using cameras and citizen scientists, published in 2015, showed that European pine martens were distributed across all counties of Northern Ireland. Their reintroduction has been controversial.


Behaviour and ecology

The European pine marten is the only
mustelid The Mustelidae (; from Latin , weasel) are a diverse family of carnivoran mammals, including weasels, badgers, otters, polecats, martens, grisons, and wolverines. Otherwise known as mustelids (), they form the largest family in the suborde ...
with semiretractable claws. This enables them to lead more
arboreal Arboreal locomotion is the locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally (scansorial), but others are exclusively arboreal. The hab ...
lifestyles, such as climbing or running on tree branches, although they are also relatively quick runners on the ground. They are mainly active at night and dusk. They have small, rounded, highly sensitive ears and sharp teeth adapted for eating small mammals, birds, insects, frogs, and
carrion Carrion (), also known as a carcass, is the decaying flesh of dead animals. Overview Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures ...
. They have also been known to eat
berries A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone fruit, stone or pit (fruit), pit although many wikt:pip#Etymology 2, pips or seeds may be p ...
, fruits, birds' eggs, nuts, and honey. The European pine marten is
territorial A territory is an area of land, sea, or space, belonging or connected to a particular country, person, or animal. In international politics, a territory is usually a geographic area which has not been granted the powers of self-government, ...
and marks its
home range A home range is the area in which an animal lives and moves on a periodic basis. It is related to the concept of an animal's territory which is the area that is actively defended. The concept of a home range was introduced by W. H. Burt in 1943. ...
by depositing
feces Feces (also known as faeces American and British English spelling differences#ae and oe, or fæces; : faex) are the solid or semi-solid remains of food that was not digested in the small intestine, and has been broken down by bacteria in the ...
. Its scat is black and twisted. The recovery of the European pine marten in Ireland has been credited with reducing the population of invasive grey squirrels. Where the range of the expanding European pine marten population meets that of the grey squirrel, the population of the grey squirrels quickly retreats and the red squirrel population recovers. Because the grey squirrel spends more time on the ground than the red squirrel, which co-evolved with the pine marten, they are thought to be far more likely to come in contact with this predator.


Lifespan

The European pine marten has lived to 18 years in captivity, but the maximum age recorded in the wild is only 11 years, with 3–4 years being more typical. They reach sexual maturity at 2–3 years of age. Copulation usually occurs on the ground and can last more than an hour. Mating occurs in July and August, but the fertilized egg does not enter the uterus for about 7 months. The young are usually born in late March or early April, in litters of one to five, after a month-long
gestation Gestation is the period of development during the carrying of an embryo, and later fetus, inside viviparous animals (the embryo develops within the parent). It is typical for mammals, but also occurs for some non-mammals. Mammals during pregn ...
period that happens after the implantation of the fertilized egg. Young European pine martens weigh around at birth. The young begin to emerge from their dens around 7–8 weeks after birth and are able to disperse from the den around 12–16 weeks after their birth.


Predators

Larger mammalian predators and birds of prey such as
Eurasian lynx The Eurasian lynx (''Lynx lynx'') is one of the four wikt:extant, extant species within the medium-sized wild Felidae, cat genus ''Lynx''. It is widely distributed from Northern Europe, Northern, Central Europe, Central and Eastern Europe to Cent ...
,
wolverine The wolverine ( , ; ''Gulo gulo''), also called the carcajou or quickhatch (from East Cree, ''kwiihkwahaacheew''), is the largest land-dwelling species, member of the family Mustelidae. It is a muscular carnivore and a solitary animal. The w ...
,
red fox The red fox (''Vulpes vulpes'') is the largest of the true foxes and one of the most widely distributed members of the order Carnivora, being present across the entire Northern Hemisphere including most of North America, Europe and Asia, plus ...
,
golden eagle The golden eagle (''Aquila chrysaetos'') is a bird of prey living in the Northern Hemisphere. It is the most widely distributed species of eagle. Like all eagles, it belongs to the family Accipitridae. They are one of the best-known bird of pr ...
, white-tailed eagle, northern goshawk, and Eurasian eagle owl prey on European pine martens, especially young individuals.


Threats

Humans are the greatest threat to the European pine marten. They are vulnerable to conflict with humans arising from predator control for other species, or following predation of livestock and the use of inhabited buildings for denning. Martens may also be affected by woodland loss, which results in habitat loss for the animal. Persecution (illegal poisoning and shooting), loss of habitat leading to fragmentation, and other human disturbances have caused a considerable decline in the European pine marten population. In some areas, they are also prized for their very fine fur. In the UK, European pine martens and their dens are offered full protection under the
Wildlife and Countryside Act 1981 The Wildlife and Countryside Act 1981 (c. 69) is an act of Parliament in the United Kingdom implemented to comply with European Council Directive 79/409/EEC on the conservation of wild birds. In short, the act gives protection to native species ...
and the Environmental Protection Act 1990.


In culture

The European pine marten is one of the national symbols of Croatia and appears on the Croatian euro coins. Previously, the European pine marten appeared on the obverse side of the 1-, 2-, 5-, and 25- Croatian kuna, the word "kuna" meaning "marten" in Croatian.


See also

* '' Marturina''


References


Further reading

* {{Authority control Martens Mammals of Azerbaijan Carnivorans of Europe Mammals described in 1758 Taxa named by Carl Linnaeus Habitats Directive species