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''Panthera gombaszoegensis'', also known as the European jaguar, is a ''
Panthera ''Panthera'' is a genus within the family (biology), family Felidae that was named and described by Lorenz Oken in 1816 who placed all the spotted cats in this group. Reginald Innes Pocock revised the classification of this genus in 1916 as co ...
''
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
that lived from about 2.0 to 0.35 million years ago in
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
. The first fossils were excavated in 1938 in Gombasek,
Slovakia Slovakia (; sk, Slovensko ), officially the Slovak Republic ( sk, Slovenská republika, links=no ), is a landlocked country in Central Europe. It is bordered by Poland to the north, Ukraine to the east, Hungary to the south, Austria to the s ...
. More of its fossil remains were excavated from the
Olivola Olivola ( pms, Aulìvola) is a ''comune'' (municipality) in the Province of Alessandria in the Italian region Piedmont, about east of Turin and about northwest of Alessandria. Olivola borders the municipalities of Casorzo, Frassinello Monferrato ...
site in
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...
, while finds elsewhere in Italy were initially named ''Panthera toscana''. Later fossils were found in
England England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe b ...
,
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
,
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
,
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
, and the
Netherlands ) , anthem = ( en, "William of Nassau") , image_map = , map_caption = , subdivision_type = Sovereign state , subdivision_name = Kingdom of the Netherlands , established_title = Before independence , established_date = Spanish Netherl ...
. Sometimes it is recognized as a subspecies of ''Panthera onca'', the
jaguar The jaguar (''Panthera onca'') is a large cat species and the only living member of the genus '' Panthera'' native to the Americas. With a body length of up to and a weight of up to , it is the largest cat species in the Americas and the th ...
, as ''Panthera onca gombaszoegensis.''


Description

European jaguars were larger than modern-day jaguars found in
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
, such as
Pantanal The Pantanal () is a natural region encompassing the world's largest tropical wetland area, and the world's largest flooded grasslands. It is located mostly within the Brazilian state of Mato Grosso do Sul, but it extends into Mato Grosso and p ...
and
Peruvian jaguar The South American jaguar is a jaguar (''Panthera onca'') population in South America. Though a number of subspecies of jaguar have been proposed for South America, morphological and genetic research did not reveal any evidence for subspecifi ...
s. With a bodyweight between 70 and 210 kg (154 to 463 lbs), they were therefore probably capable of bringing down larger prey. Like other ''Panthera'' species, they are thought to have been sexually dimorphic, with significantly larger males.


History

The ancestors of jaguars are thought to have arisen in Africa; a related form of ''Panthera'' was present in South Africa 1.9 Ma ago. Another form similar to ''P. gombaszoegensis'' has been found dating from early
Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...
East Africa East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territories make up Eastern Africa: Due to the historical ...
and had both
lion The lion (''Panthera leo'') is a large Felidae, cat of the genus ''Panthera'' native to Africa and India. It has a muscular, broad-chested body; short, rounded head; round ears; and a hairy tuft at the end of its tail. It is sexually dimorphi ...
- and
tiger The tiger (''Panthera tigris'') is the largest living cat species and a member of the genus '' Panthera''. It is most recognisable for its dark vertical stripes on orange fur with a white underside. An apex predator, it primarily preys on u ...
-like characteristics. ''P. gombaszoegensis'' was initially the only European
Pantherinae Pantherinae is a subfamily within the family Felidae; it was named and first described by Reginald Innes Pocock in 1917 as only including the ''Panthera'' species. The Pantherinae genetically diverged from a common ancestor between and . Char ...
species in the Early Pleistocene, being present alongside the felines ''
Acinonyx pardinensis The giant cheetah (''Acinonyx pardinensis'') is an extinct felid species that was closely related to the modern cheetah. Description The lifestyle and physical characteristics of the giant cheetah were probably similar to those of its modern ...
'' and ''
Puma pardoides ''Puma pardoides'', sometimes called the Eurasian puma or Owen's panther, is an extinct prehistoric cat. It was long regarded as a primitive species of leopard (genus ''Panthera''). Recent work however has shown that ''Panthera pardoides'' and '' ...
'' and the machairodontines ''
Homotherium ''Homotherium'', also known as the scimitar-toothed cat or scimitar cat, is an extinct genus of machairodontine saber-toothed predator, often termed scimitar-toothed cats, that inhabited North America, South America, Eurasia, and Africa during th ...
latidens'' and ''
Megantereon ''Megantereon'' was a genus of prehistoric machairodontine saber-toothed cat that lived in North America, Eurasia, and Africa. It may have been the ancestor of ''Smilodon''. Taxonomy Fossil fragments have been found in Africa, Eurasia, and No ...
whitei''.
Leopard The leopard (''Panthera pardus'') is one of the five extant species in the genus '' Panthera'', a member of the cat family, Felidae. It occurs in a wide range in sub-Saharan Africa, in some parts of Western and Central Asia, Southern Russia, a ...
s arrived later in the Early Pleistocene or the Middle Pleistocene, and
lions The lion (''Panthera leo'') is a large cat of the genus ''Panthera'' native to Africa and India. It has a muscular, broad-chested body; short, rounded head; round ears; and a hairy tuft at the end of its tail. It is sexually dimorphic; adult ...
in the Middle Pleistocene.


Taxonomy

''Leo gombaszoegensis'' was the
scientific name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
proposed by
Miklós Kretzoi Miklós Kretzoi (9 February 1907 – 15 March 2005) was a Hungarian geologist, paleontologist and paleoanthropologist and Széchenyi Prize winner. Exhibition in the Hungarian National Museum, 9 February - 24 May 2004 Kretzoi studied A ...
in 1938 for teeth found in
Tertiary Tertiary ( ) is a widely used but obsolete term for the geologic period from 66 million to 2.6 million years ago. The period began with the demise of the non-avian dinosaurs in the Cretaceous–Paleogene extinction event, at the start ...
deposits in
Gombasek Cave Gombasek Cave or Gombasecká jaskyňa (in Slovak language, Slovak) and Gombaszögi-barlang (in Hungarian language, Hungarian) is a karst cave in the Slovak Karst, Slovakia. It is named after the settlement of ''Gombasek'', which belongs to the ...
,
Slovakia Slovakia (; sk, Slovensko ), officially the Slovak Republic ( sk, Slovenská republika, links=no ), is a landlocked country in Central Europe. It is bordered by Poland to the north, Ukraine to the east, Hungary to the south, Austria to the s ...
. It was reassessed and subordinated to the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''Panthera'' in 1971. The following are considered to belong to ''P. gombaszoegensis'' as well: *''Panthera toscana'' proposed in 1949 for
carnassial Carnassials are paired upper and lower teeth modified in such a way as to allow enlarged and often self-sharpening edges to pass by each other in a shearing manner. This adaptation is found in carnivorans, where the carnassials are the modified f ...
teeth found in
Villafranchian Villafranchian age ( ) is a period of geologic time (3.5–1.0 Ma) spanning the Late Pliocene and Early Pleistocene used more specifically with European Land Mammal Ages. Named by Italian geologist Lorenzo Pareto for a sequence of terrestrial s ...
deposits in the Arno river valley in Italy. These remains were originally described as a distinct species and later as subspecies ''Panthera gombaszoegensis toscana''. It is sometimes referred to as the Tuscany jaguar or Tuscany lion. *''Felis (Panthera) schreuderi'' proposed in 1960 for cat fossils found in
Tegelen Tegelen ( li, Tegele) is a village in the municipality of Venlo, situated in the Netherlands. It was an independent municipality until 2001, when it was merged into the municipality of Venlo. Tiglian The name of the glacial stage of Tiglian (pa ...
, the Netherlands. *''Jansofelis vaufreyi'' proposed in 1971 for cat fossils found in southeastern France. Some remains once attributed to ''P. gombaszoegensis'' have more recently been identified as ''
Acinonyx pardinensis The giant cheetah (''Acinonyx pardinensis'') is an extinct felid species that was closely related to the modern cheetah. Description The lifestyle and physical characteristics of the giant cheetah were probably similar to those of its modern ...
''. A 2022 study based on a relatively complete skull from Belgium suggested that ''P. gombaszoegensis'' is more closely related to the
tiger The tiger (''Panthera tigris'') is the largest living cat species and a member of the genus '' Panthera''. It is most recognisable for its dark vertical stripes on orange fur with a white underside. An apex predator, it primarily preys on u ...
(''Panthera tigris''), than the jaguar.


Habitat and behavior

The European jaguar has often been thought to be a forest-dwelling
cat The cat (''Felis catus'') is a domestic species of small carnivorous mammal. It is the only domesticated species in the family Felidae and is commonly referred to as the domestic cat or house cat to distinguish it from the wild members of ...
, similar in habits to the modern
jaguar The jaguar (''Panthera onca'') is a large cat species and the only living member of the genus '' Panthera'' native to the Americas. With a body length of up to and a weight of up to , it is the largest cat species in the Americas and the th ...
, although recent work suggests that the association between the European jaguar and forested habitats is not as strong as has often been assumed. It was probably a
solitary Solitary is the state of being alone or in solitude. The term may refer to: * shortened form of solitary confinement * Solitary animal, an animal that does not live with others in its species * Solitary but social, a type of social organization in ...
animal. File:Panthera toscana.JPG, Skull,
Florence Florence ( ; it, Firenze ) is a city in Central Italy and the capital city of the Tuscany region. It is the most populated city in Tuscany, with 383,083 inhabitants in 2016, and over 1,520,000 in its metropolitan area.Bilancio demografico an ...
File:Panthera toscana 1.JPG, Same skull File:Panthera schreuderi.JPG, Mandible,
Rotterdam Rotterdam ( , , , lit. ''The Dam on the River Rotte'') is the second largest city and municipality in the Netherlands. It is in the province of South Holland, part of the North Sea mouth of the Rhine–Meuse–Scheldt delta, via the ''"N ...


See also

*''
Panthera atrox ''Panthera atrox'', better known as the American lion, also called the North American lion, or American cave lion, is an extinct Pantherinae, pantherine cat that lived in North America during the Pleistocene Epoch (geology), epoch and the earl ...
'' *''
Panthera blytheae ''Panthera blytheae'' is an extinct species of the genus '' Panthera'' that is thought to have existed during the late Messinian to early Zanclean ages approximately 5.95–4.1 million years ago. It is currently the oldest known ''Panthera'' sp ...
'' *''
Panthera palaeosinensis ''Panthera palaeosinensis'' was an early Pleistocene species from northern China. It is often incorrectly referenced as the ancestor of the tiger, ''Panthera tigris'', although it shares features with all living large cats. Recent studies place i ...
'' *'' Panthera shawi'' *''
Panthera spelaea ''Panthera spelaea'', also known as the Eurasian cave lion, European cave lion or steppe lion, is an extinct ''Panthera'' species that most likely evolved in Europe after the third Cromerian interglacial stage, less than 600,000 years ago. Phylo ...
'' *''
Panthera zdanskyi ''Panthera zdanskyi'', also known as the Longdan tiger, is an extinct pantherine species that is seen as a close relative of the modern tiger. Fossils were excavated in northwestern China's Gansu province. Etymology ''Panthera zdanskyi'' was fi ...
''


References

{{Taxonbar, from=Q1164873 Panthera Pleistocene carnivorans Pleistocene mammals of Europe Pleistocene mammals of Africa Middle Pleistocene Prehistoric pantherines Prehistoric mammals of Europe Prehistoric mammals of Africa Fossil taxa described in 1938 Pleistocene first appearances Pleistocene extinctions