The European wild ass (''Equus hemionus hydruntinus'') or hydruntine is an extinct
equine
Equinae is a subfamily of the family Equidae, which have lived worldwide (except Indonesia and Australia) from the Hemingfordian stage of the Early Miocene (16 million years ago) onwards. They are thought to be a monophyletic grouping.B. J. MacFa ...
from the
Middle Pleistocene
The Chibanian, widely known by its previous designation of Middle Pleistocene, is an age in the international geologic timescale or a stage in chronostratigraphy, being a division of the Pleistocene Epoch within the ongoing Quaternary Period. The ...
to Late
Holocene
The Holocene ( ) is the current geological epoch. It began approximately 11,650 cal years Before Present (), after the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene togethe ...
of Western Eurasia. It appeared first in the fossil record around 350–300,000 years before present. In the late Pleistocene it was widespread throughout much of western Eurasia from the Middle East to Europe, especially along the
Mediterranean
The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on the e ...
, with fossil reports from
Sicily
(man) it, Siciliana (woman)
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, population_blank1 =
, demographics_type1 = Ethnicity
, demographics1_footnotes =
, demographi ...
, Turkey, Spain, France and Portugal. In the east the range apparently stretched at least to the Volga and to Iran. In the north it reached almost to the
North Sea
The North Sea lies between Great Britain, Norway, Denmark, Germany, the Netherlands and Belgium. An epeiric sea on the European continental shelf, it connects to the Atlantic Ocean through the English Channel in the south and the Norwegian S ...
in Germany and the
British Isles
The British Isles are a group of islands in the North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner and Outer Hebrides, the Northern Isles, ...
.
Its range fragmented after the
Last Glacial Maximum
The Last Glacial Maximum (LGM), also referred to as the Late Glacial Maximum, was the most recent time during the Last Glacial Period that ice sheets were at their greatest extent.
Ice sheets covered much of Northern North America, Northern Eur ...
, surviving into the Holocene, its range gradually contracted further, persisting in small regions of
southern Europe
Southern Europe is the southern regions of Europe, region of Europe. It is also known as Mediterranean Europe, as its geography is essentially marked by the Mediterranean Sea. Definitions of Southern Europe include some or all of these countrie ...
into the
Bronze Age
The Bronze Age is a historic period, lasting approximately from 3300 BC to 1200 BC, characterized by the use of bronze, the presence of writing in some areas, and other early features of urban civilization. The Bronze Age is the second pri ...
, and in Iran and the
Caucasus
The Caucasus () or Caucasia (), is a region between the Black Sea and the Caspian Sea, mainly comprising Armenia, Azerbaijan, Georgia, and parts of Southern Russia. The Caucasus Mountains, including the Greater Caucasus range, have historically ...
into the
Iron Age
The Iron Age is the final epoch of the three-age division of the prehistory and protohistory of humanity. It was preceded by the Stone Age (Paleolithic, Mesolithic, Neolithic) and the Bronze Age (Chalcolithic). The concept has been mostly appl ...
, around 500 BC.
It has been suggested that the Iberian Zebro, extinct in the wild from the 16th century, could correspond to the ''Equus hydruntinus'', although the word "
zebro" or "cebro" comes from Latin ''equiferus'' meaning 'wild horse'. Later research judged that it was unlikely that hydruntines persisted in the
Iberian Peninsula
The Iberian Peninsula (),
**
* Aragonese and Occitan: ''Peninsula Iberica''
**
**
* french: Péninsule Ibérique
* mwl, Península Eibérica
* eu, Iberiar penintsula also known as Iberia, is a peninsula in southwestern Europe, defi ...
beyond the
Chalcolithic
The Copper Age, also called the Chalcolithic (; from grc-gre, χαλκός ''khalkós'', "copper" and ''líthos'', "stone") or (A)eneolithic (from Latin '' aeneus'' "of copper"), is an archaeological period characterized by regular ...
.
Morphologically the European ass can be distinguished from
asses
Ass most commonly refers to:
* Buttocks (in informal American English)
* Donkey or ass, ''Equus africanus asinus''
**any other member of the subgenus ''Asinus''
Ass or ASS may also refer to:
Art and entertainment
* Ass (album), ''Ass'' (album ...
and
hemiones particularly by its
molars
The molars or molar teeth are large, flat teeth at the back of the mouth. They are more developed in mammals. They are used primarily to grind food during chewing. The name ''molar'' derives from Latin, ''molaris dens'', meaning "millstone to ...
and the relatively short
nare
A nostril (or naris , plural ''nares'' ) is either of the two orifices of the nose. They enable the entry and exit of air and other gasses through the nasal cavities. In birds and mammals, they contain branched bones or cartilages called tur ...
s. The exact systematic position was formerly unclear but genetic and morphological analysis suggested that it is closely related to the
Asiatic wild ass. A 2017 genetic study found that it was a subspecies of Asiatic wild ass, closer to the
Khur
The Indian wild ass (''Equus hemionus khur''), also called the Indian onager or, in the local Gujarati language, Ghudkhur and Khur, is a subspecies of the onager native to South Asia.
It is currently listed as Near Threatened by IUCN. The pr ...
than the
Persian onager
The Persian onager (''Equus hemionus onager''), also called the Persian wild ass or Persian zebra, is a subspecies of onager (Asiatic wild ass) native to Iran (Persia). It is listed as Endangered, with no more than 600 individuals left in the wil ...
.
Etymology
The specific epithet, ''hydruntinus'', means ''from
Otranto
Otranto (, , ; scn, label= Salentino, Oṭṛàntu; el, label=Griko, Δερεντό, Derentò; grc, Ὑδροῦς, translit=Hudroûs; la, Hydruntum) is a coastal town, port and ''comune'' in the province of Lecce (Apulia, Italy), in a fertil ...
'' (''
Hydruntum
Otranto (, , ; scn, label= Salentino, Oṭṛàntu; el, label=Griko, Δερεντό, Derentò; grc, Ὑδροῦς, translit=Hudroûs; la, Hydruntum) is a coastal town, port and ''comune'' in the province of Lecce ( Apulia, Italy), in a ferti ...
'' in
Latin
Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
).
Ecology
The evidence shows that the European ass favoured semi-arid,
steppic conditions and showed a preference for
temperate
In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout t ...
climates, although it was also found in cool or cold conditions.
It may have retreated to warmer locales during the coldest periods
although the relatively short muzzle indicates an adaptation for cold conditions. It preferred open
biotopes, between
shrubland
Shrubland, scrubland, scrub, brush, or bush is a plant community characterized by vegetation dominated by shrubs, often also including grasses, herbs, and geophytes. Shrubland may either occur naturally or be the result of human activity. It m ...
(favoured by
true horses) and
grassland
A grassland is an area where the vegetation is dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur natur ...
(favoured by
bovids
The Bovidae comprise the biological family of cloven-hoofed, ruminant mammals that includes cattle, bison, buffalo, antelopes, and caprines. A member of this family is called a bovid. With 143 extant species and 300 known extinct species, the ...
). It is believed to have shared this habitat with species such as the
woolly rhinoceros
The woolly rhinoceros (''Coelodonta antiquitatis'') is an extinct species of rhinoceros that was common throughout Europe and Asia during the Pleistocene epoch and survived until the end of the last glacial period. The woolly rhinoceros was a me ...
.
It is considered an ecologically important part of the ecosystem known as
mammoth steppe
During the Last Glacial Maximum, the mammoth steppe, also known as steppe-tundra, was the Earth's most extensive biome. It spanned from Spain eastward across Eurasia to Canada and from the List of islands in the Arctic Ocean, arctic islands sout ...
where it filled a niche equivalent to that provided by the
African wild ass
The African wild ass (''Equus africanus'') or African wild donkey is a wild member of the horse family, Equidae. This species is thought to be the ancestor of the domestic donkey (''Equus asinus''), which is sometimes placed within the same spec ...
or
Zebra
Zebras (, ) (subgenus ''Hippotigris'') are African equines with distinctive black-and-white striped coats. There are three living species: the Grévy's zebra (''Equus grevyi''), plains zebra (''E. quagga''), and the mountain zebra (''E. zeb ...
in the African
Savanna
A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground to ...
.
See also
*
List of extinct animals of Europe
This list of European animals extinct in the Holocene features animals known to have become extinct in the last 12,000 years on the European continent and its surrounding islands.
Dependent territories of European countries in other continents, l ...
References
Onager
Pleistocene mammals of Europe
Extinct mammals of Europe
Holocene extinctions
Middle Pleistocene
Late Pleistocene
Extinct mammals
Mammals described in 1907
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