Eucalyptus Botryoides
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''Eucalyptus botryoides'', commonly known as the bangalay, bastard jarrah, woollybutt or southern mahogany, is a small to tall tree native to southeastern Australia. Reaching up to high, it has rough bark on its trunk and branches. It is found on sandstone- or shale-based soils in open woodland, or on more sandy soils behind sand dunes. The white flowers appear in summer and autumn. It reproduces by resprouting from its woody
lignotuber A lignotuber is a woody swelling of the root crown possessed by some plants as a protection against destruction of the plant stem, such as by fire. Other woody plants may develop basal burls as a similar survival strategy, often as a response t ...
or epicormic buds after bushfire. ''E. botryoides'' hybridises with the Sydney blue gum ('' E. saligna'') in the Sydney region. The hard, durable wood has been used for panelling and flooring.


Description

In favourable conditions, ''Eucalyptus botryoides'' can grow as a straight-trunked tree to high with a dbh of , although it is often shorter in poorer situations. In exposed areas behind sand dunes, it is a lower spreading tree tall, with its leaves forming a dense
crown A crown is a traditional form of head adornment, or hat, worn by monarchs as a symbol of their power and dignity. A crown is often, by extension, a symbol of the monarch's government or items endorsed by it. The word itself is used, partic ...
, or even a multitrunked mallee form in poor sandy soils. It has a swollen woody base known as a
lignotuber A lignotuber is a woody swelling of the root crown possessed by some plants as a protection against destruction of the plant stem, such as by fire. Other woody plants may develop basal burls as a similar survival strategy, often as a response t ...
which can reach in diameter.Benson and McDougall (1998), p. 871. The thick, fibrous, rough and flaky bark covers the trunk and larger branches, and is vertically furrowed. Retrieved 19 January 2012. The bark is more greyish brown in trees of inland forest origin, and a redder brown in those of more coastal origin. The bark on smaller branches is smooth and pale grey. The adult leaves are stalked, broad-lanceolate, to long by wide, and are dark green above, and paler below. Venation is fine and at 40° – 60° to the midline. Developing from small cylindrical or club-shaped (clavate) buds, the white flowers appear from January to April, and are arranged in groups of six to eleven in umbellasters. The woody fruits, or gumnuts, are ovoid or cylindrical in shape, and measure between 7–12 mm long and 4–6 mm wide, with the valve near the rim or enclosed. Seedlings and young plants have more ovate leaves which are arranged oppositely along the stems for the first three to six pairs until they assume the adult alternately arranged configuration. They are also paler on the undersurface, and measure 4.5 to 11 cm long and 1.3 to 5.5 cm wide.


Taxonomy

The tree was first described by naturalist
James Edward Smith James Edward Smith may refer to: * James Edward Smith (botanist), English botanist and founder of the Linnean Society * James Edward Smith (murderer), American murderer * James Edward Smith (politician), Canadian businessman and mayor of Toronto * ...
in 1797, without nominating a type specimen, and still bears its original name. The species name is derived from the
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic peri ...
''botrys'' "cluster", and may relate to the clustered flowerheads and fruit. It has been classified in the subgenus ''Symphyomyrtus'', Section ''Latoangulatae'', Series ''Annulares'' (red mahoganies) by Brooker and Kleinig. Its closest relatives are the red mahogany ('' Eucalyptus scias'') and the Blue Mountains mahogany ('' E. notabilis''), red mahogany/red messmate ('' E. resinifera'') and swamp mahogany ('' E. robusta''). South of Sydney Harbour and Parramatta River, ''E. botryoides'' forms hybrid populations with Sydney blue gum ('' E. saligna'').


Distribution and habitat

Distribution is coastal south eastern Australia from near
Newcastle Newcastle usually refers to: *Newcastle upon Tyne, a city and metropolitan borough in Tyne and Wear, England *Newcastle-under-Lyme, a town in Staffordshire, England *Newcastle, New South Wales, a metropolitan area in Australia, named after Newcastle ...
on the mid coast of
New South Wales ) , nickname = , image_map = New South Wales in Australia.svg , map_caption = Location of New South Wales in AustraliaCoordinates: , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , es ...
to eastern
Victoria Victoria most commonly refers to: * Victoria (Australia), a state of the Commonwealth of Australia * Victoria, British Columbia, provincial capital of British Columbia, Canada * Victoria (mythology), Roman goddess of Victory * Victoria, Seychelle ...
in the
Lakes Entrance Lakes Entrance is a seaside resort and fishing port in eastern Victoria, Australia. It is situated approximately east of Melbourne, near a managed, artificial channel connecting the Gippsland Lakes to Bass Strait. At the 2016 census, Lakes Ent ...
area, specifically Loch Sport south of
Bairnsdale Bairnsdale () ( Ganai: ''Wy-yung'') is a city in East Gippsland, Victoria, Australia in a region traditionally owned by the Tatungalung clan of the Gunaikurnai people. The estimated population of Bairnsdale urban area was 15,411 at Ju ...
. The species was introduced to Western Australia, where it is listed as an
alien Alien primarily refers to: * Alien (law), a person in a country who is not a national of that country ** Enemy alien, the above in times of war * Extraterrestrial life, life which does not originate from Earth ** Specifically, intelligent extrater ...
. It grows predominantly on low nutrient sandstone-derived or sandy soils, either behind coastal sand dunes or further inland in alluvial soils in valleys, where it is a dominant tree. It is generally not far from salt water. ''Eucalyptus botryoides'' is only found in lowlands, from sea level to altitude, and in areas of rainfall from . Trees in mixed open forest it grows with include turpentine (''
Syncarpia glomulifera ''Syncarpia glomulifera'', commonly known as the turpentine tree, or yanderra, is a tree of the family Myrtaceae native to New South Wales and Queensland in Australia, which can reach in height. It generally grows on heavier soils. The cream fl ...
''), spotted gum (''
Corymbia maculata ''Corymbia maculata'', commonly known as spotted gum, is a species of medium-sized to tall tree that is endemic to eastern Australia. It has smooth, mottled bark, lance-shaped to curved adult leaves, flower buds usually in groups of three, whit ...
''), red bloodwood ('' C. gummifera''), blackbutt ('' E. pilularis''), Sydney blue gum, red mahogany ('' E. resinifera''), and smooth-barked apple (''
Angophora costata ''Angophora costata'', commonly known as Sydney red gum, rusty gum or smooth-barked apple, is a species of tree that is endemic to eastern Australia. Reaching in height, the species has distinctive smooth bark that is pinkish or orange-brown wh ...
''). Associated
understory In forestry and ecology, understory (American English), or understorey (Commonwealth English), also known as underbrush or undergrowth, includes plant life growing beneath the forest canopy without penetrating it to any great extent, but abov ...
plants in wetter forests with some rainforest transition include lillypilly (''
Syzygium smithii ''Syzygium smithii'' (formerly ''Acmena smithii'') is a summer-flowering, winter-fruiting evergreen tree, belonging to the myrtle family Myrtaceae. It shares the common name "lilly pilly" with several other plants.It is planted as shrubs or hed ...
'') and wattles. In coastal plant communities near sand-dunes, it grows with stunted forms of white stringybark ('' E. globoidea''), silvertop ash ('' E. sieberi''),
banksia ''Banksia'' is a genus of around 170 species in the plant family Proteaceae. These Australian wildflowers and popular garden plants are easily recognised by their characteristic flower spikes, and fruiting "cones" and heads. ''Banksias'' range i ...
and such understory plants as burrawang (''
Macrozamia communis ''Macrozamia communis'' is an Australian cycad found on the east coast of New South Wales. The common name for the species is burrawang, a word derived from the Daruk Australian Aboriginal language; this name is also often applied to other spec ...
''). It is a component tree of wetland forest in
Booderee National Park Booderee National Park and Botanic Gardens, formerly Jervis Bay National Park and Jervis Bay Botanic Gardens, are located in the Jervis Bay Territory of Australia. The reserve is composed of two sections: * the Bherwerre Peninsula, on the souther ...
alongside blackbutt, red bloodwood, grey ironbark (''
Eucalyptus paniculata ''Eucalyptus paniculata'', commonly known as grey ironbark, is a species of tree that is endemic to New South Wales. It has dark-coloured, deeply furrowed ironbark on the trunk and branches, lance-shaped to curved adult leaves, flower buds in gro ...
''), scribbly gum ('' E. sclerophylla''), old man banksia (''
Banksia serrata ''Banksia serrata'', commonly known as the saw banksia, the old man banksia, the saw-tooth banksia or the red honeysuckle and as wiriyagan by the Cadigal people, is a species of woody shrub or tree of the genus ''Banksia'', in the family Prote ...
''), coast banksia ('' B. integrifolia'') and snow-in-summer (''
Melaleuca linariifolia ''Melaleuca linariifolia'' is a plant in the myrtle family Myrtaceae, and is endemic to eastern Australia. It is commonly known as snow-in-summer, narrow-leaved paperbark, flax-leaved paperbark and in the language of the Gadigal people as ''budj ...
''), with understory plants such as jointed twig-rush (''
Baumea articulata ''Baumea articulata'', commonly known as jointed rush, is a sedge in the sedge family, Cyperaceae, that is native to Western Australia. The grass-like plant is rhizomatous and perennial, it typically grows to a height of . It blooms between Septe ...
''), tall spike-rush ('' Eleocharis sphacelata''), prickly tea-tree ('' Leptospermum juniperinum''), and zig-zag bog-rush (''
Schoenus brevifolius ''Schoenus brevifolius'', known as zig-zag bog-rush, is a species of sedge native to Australia, New Zealand, New Caledonia, and the Ogasawara ( Bonin) Islands. It was first described by Robert Brown in 1810.Govaerts, R. & Simpson, D.A. (2007 ...
'').


Ecology

''Eucalyptus botryoides'' regenerates after bushfire by resprouting from epicormic buds and its woody lignotuber. Plants have been dated at 600 years of age. It also drops branches, and these have been known to grow roots. The wet environment and water-absorbing qualities of the thick, fibrous bark facilitate this. The
koala The koala or, inaccurately, koala bear (''Phascolarctos cinereus''), is an arboreal herbivorous marsupial native to Australia. It is the only extant representative of the family Phascolarctidae and its closest living relatives are the womb ...
(''Phascalarctos cinereus'') eats the leaves, and ants consume the nectar. The species is highly susceptible to psyllids.


Uses

The heartwood of this species is durable and heavy (about 765–985 kg/m3), and resembles that of ''E. saligna'' and ''E. grandis''. It is used for flooring and panelling. File:Bangalay-bark.jpg, Trunk bark File:Bangalay-leaves.jpg, Adult leaves File:Bangalay-leaves-juvenile.jpg, Juvenile leaves File:Bangalay-fruit.jpg, Buds File:Bangalay-flowers.jpg, Inflorescence File:Bangalay-flowering.jpg, Flowering tree File:Bangalay-bird-hollow.jpg, Bird hollow


References

{{Taxonbar, from=Q2484889 botryoides Myrtales of Australia Trees of Australia Flora of Victoria (Australia) Flora of New South Wales