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The Ethiopian hare (''Lepus fagani'') is a species of
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
in the family
Leporidae Leporidae is the family of rabbits and hares, containing over 60 species of extant mammals in all. The Latin word ''Leporidae'' means "those that resemble ''lepus''" (hare). Together with the pikas, the Leporidae constitute the mammalian order ...
. It was first described in 1903, by the British
mammalogist In zoology, mammalogy is the study of mammals – a class of vertebrates with characteristics such as homeothermic metabolism, fur, four-chambered hearts, and complex nervous system In biology, the nervous system is the highly complex part ...
Oldfield Thomas Michael Rogers Oldfield Thomas (21 February 1858 – 16 June 1929) was a British zoologist. Career Thomas worked at the Natural History Museum on mammals, describing about 2,000 new species and subspecies for the first time. He was appoin ...
. The dorsal
pelage Fur is a thick growth of hair that covers the skin of mammals. It consists of a combination of oily #Guard hair, guard hair on top and thick #Down hair, underfur beneath. The guard hair keeps moisture from reaching the skin; the underfur acts as ...
is brownish buff, and is finely grizzled with black. The ventral pelage is fluffy and white in colour.
Endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to
Ethiopia Ethiopia, , om, Itiyoophiyaa, so, Itoobiya, ti, ኢትዮጵያ, Ítiyop'iya, aa, Itiyoppiya officially the Federal Democratic Republic of Ethiopia, is a landlocked country in the Horn of Africa. It shares borders with Eritrea to the ...
, it is found in the Afromontane Biozone of Ethiopia, and in the borders of the Sudanian Savanna Biozone. It is rated as a
least concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. T ...
 species by the 
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
.


Taxonomy

The Ethiopian hare is a species of the family
Leporidae Leporidae is the family of rabbits and hares, containing over 60 species of extant mammals in all. The Latin word ''Leporidae'' means "those that resemble ''lepus''" (hare). Together with the pikas, the Leporidae constitute the mammalian order ...
, and has no recognized subspecies. It belongs to the subgenus ''Sabanalagus''. It was first described in 1903 by the British
mammalogist In zoology, mammalogy is the study of mammals – a class of vertebrates with characteristics such as homeothermic metabolism, fur, four-chambered hearts, and complex nervous system In biology, the nervous system is the highly complex part ...
Oldfield Thomas Michael Rogers Oldfield Thomas (21 February 1858 – 16 June 1929) was a British zoologist. Career Thomas worked at the Natural History Museum on mammals, describing about 2,000 new species and subspecies for the first time. He was appoin ...
in the ''
Proceedings of the Zoological Society of London The ''Journal of Zoology'' is a scientific journal concerning zoology, the study of animals. It was founded in 1830 by the Zoological Society of London and is published by Wiley-Blackwell. It carries original research papers, which are targeted ...
''. He named it ''Lepus fagani'', after Charles E. Fagan, who was the Assistant Secretary of the
British Museum of Natural History The Natural History Museum in London is a museum that exhibits a vast range of specimens from various segments of natural history. It is one of three major museums on Exhibition Road in South Kensington, the others being the Science Museum an ...
. Describing the hare's taxonomy in 1986,
Derek Yalden Derek William Yalden (4 November 1940 – 5 February 2013) was an eminent British zoologist and academic. He was an Honorary Reader at the University of Manchester. After obtaining a 1st Class B.Sc. University College London in 1962, he compl ...
and colleagues wrote "its nomenclatural history has been very confused". In 1959 and 1964 respectively, and A. A. Gureev classified it as a subspecies of ''L. crawhayi'', now regarded as a junior synonym of ''L. microtis'', the African savanna hare. In 1972, and
Derek Yalden Derek William Yalden (4 November 1940 – 5 February 2013) was an eminent British zoologist and academic. He was an Honorary Reader at the University of Manchester. After obtaining a 1st Class B.Sc. University College London in 1962, he compl ...
suggested it might be a synonym of ''L. whytei'', now a subspecies of ''L. microtis''. Petter then classified it as a subspecies of the Abyssinian hare ('' Lepus habessinicus''), in 1963. In 1987,
Maria Luisa Azzaroli Puccetti Maria may refer to: People * Mary, mother of Jesus * Maria (given name), a popular given name in many languages Place names Extraterrestrial *170 Maria, a Main belt S-type asteroid discovered in 1877 *Lunar maria (plural of ''mare''), large, da ...
reinstated its status as an independent species, because it has a uniform character throughout its distribution. It has been listed as a separate species in various works published since then. North of the
Rift Valley A rift valley is a linear shaped lowland between several highlands or mountain ranges created by the action of a geologic rift. Rifts are formed as a result of the pulling apart of the lithosphere due to extensional tectonics. The linear dep ...
, on the Ethiopian mountains, the population of the Ethiopian hare is geographically isolated from that of the cape hare in the surrounding
lowland Upland and lowland are conditional descriptions of a plain based on elevation above sea level. In studies of the ecology of freshwater rivers, habitats are classified as upland or lowland. Definitions Upland and lowland are portions of ...
s. Flux and Angermann suggested that this isolation in population probably lead to the evolution of the Ethiopian hare as a different species. In 1990, Flux and Angermann wrote it is "probably" a subspecies of the African savanna hare or of the scrub hare ('' Lepus saxatilis''). It is usually considered to be a part of the African savanna hare-scrub hare complex. ''The Kingdon Field Guide to African Mammals'' also treats it as a subspecies of the cape hare (''
Lepus capensis The Cape hare (''Lepus capensis''), also called the brown hare and the desert hare, is a hare native to Africa and Arabia extending into India. Taxonomy The Cape hare was one of the many mammal species originally described by Carl Linnaeus i ...
''). A genetic study suggests that the Ethiopian hare recently descended from the Abyssinian hare. Its
chromosome number Ploidy () is the number of complete sets of chromosomes in a cell, and hence the number of possible alleles for autosomal and pseudoautosomal genes. Sets of chromosomes refer to the number of maternal and paternal chromosome copies, respectively ...
is likely the same as other species of its genus. The
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
was collected by Edward Degen and its type locality is the
Zege Peninsula The Zege Peninsula is a peninsula located on the southern shore of Lake Tana in Ethiopia, the largest lake in Ethiopia and the source of the Blue Nile river, and is situated at (11° 40’ to 11° 43’ N and 37 °19’ to 37 °21’ E). It is 6 ...
,
Lake Tana Lake Tana ( am, ጣና ሐይቅ, T’ana ḥāyik’i; previously Tsana) is the largest lake in Ethiopia and the source of the Blue Nile. Located in Amhara Region in the north-western Ethiopian Highlands, the lake is approximately long and wid ...
, Ethiopia at an elevation of .


Description

The Ethiopian hare is a medium-sized, dark coloured hare, which measures in length. It has a medium-sized, long, fluffy tail which is buff-white below, black above, and buff coloured at the sides. The skull measures in length. The supraorbital wings are weak and small. The anterior shoulders of the
cheek bone In the human skull, the zygomatic bone (from grc, ζῠγόν, zugón, yoke), also called cheekbone or malar bone, is a paired irregular bone which articulates with the maxilla, the temporal bone, the sphenoid bone and the frontal bone. It is si ...
are large. It has long, dense, coarse
pelage Fur is a thick growth of hair that covers the skin of mammals. It consists of a combination of oily #Guard hair, guard hair on top and thick #Down hair, underfur beneath. The guard hair keeps moisture from reaching the skin; the underfur acts as ...
. The dorsal hairs are long. They are brownish buff, and are finely grizzled with black. The fur is pale grey at the base, and have buff terminal bands, whitish subterminal bands, and generally black tips. The
underfur Fur is a thick growth of hair that covers the skin of mammals. It consists of a combination of oily guard hair on top and thick underfur beneath. The guard hair keeps moisture from reaching the skin; the underfur acts as an insulating blanket ...
is whitish grey. The head is darker than the dorsal pelage, and the
crown A crown is a traditional form of head adornment, or hat, worn by monarchs as a symbol of their power and dignity. A crown is often, by extension, a symbol of the monarch's government or items endorsed by it. The word itself is used, partic ...
(top of the head) is blacker. The chin, belly, and throat are white. It has grey below the chin. The ventral pelage is fluffy and white in colour. The upperparts are buff ochraceous brown. The colour of the chest and the
nape The nape is the back of the neck. In technical anatomical/medical terminology, the nape is also called the nucha (from the Medieval Latin rendering of the Arabic , "spinal marrow"). The corresponding adjective is ''nuchal'', as in the term ''nu ...
(back of the neck) are tawny. The rich cinnamon-ginger colour of the
nuchal The nape is the back of the neck. In technical anatomical/medical terminology, the nape is also called the nucha (from the Medieval Latin rendering of the Arabic , "spinal marrow"). The corresponding adjective is ''nuchal'', as in the term ''nu ...
patch, extends on to the neck-sides. The ears are medium-sized, buff coloured, and measure in length. The lower outer-margins of the ears have buff or white fringe, the upper outer-margins have buff fringe, and the inner surfaces of the tip of the ears have narrow, black rims. The principal
incisor Incisors (from Latin ''incidere'', "to cut") are the front teeth present in most mammals. They are located in the premaxilla above and on the mandible below. Humans have a total of eight (two on each side, top and bottom). Opossums have 18, whe ...
teeth have deep grooves filled completely with cement. The dorsal parts of the upper incisor teeth are sloped laterally, forming an angled anterior surface. The forefeet are cinnamon-brown and have blackish brown furs at the soles. The hindfeet are white below, brownish buff above, and measure in length. The soles of the hindfeet have ginger-brown or blackish brown fur. The Abyssinian hare is similar to the Ethiopian hare, but it has soft dorsal pelage, longer ears, and a longer tail. The Ethiopian highland hare ('' Lepus starcki'') is also a similar species, but it has longer ears, a longer tail, and its grooves of the principal upper incisor teeth are not filled with cement. The cape hare is also similar to the Ethiopian hare, but has longer ears, grizzled-greyish pelage, and brownish pink nuchal patch.


Distribution and habitat

The Ethiopian hare is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
, and is found in the Afromontane Biozone of Ethiopia, and in the borders of the Sudanian Savanna Biozone; it also occurs west of the Rift Valley, in the
Ethiopian Highlands The Ethiopian Highlands is a rugged mass of mountains in Ethiopia in Northeast Africa. It forms the largest continuous area of its elevation in the continent, with little of its surface falling below , while the summits reach heights of up to . ...
, and abundantly found around the
Lake Tana Lake Tana ( am, ጣና ሐይቅ, T’ana ḥāyik’i; previously Tsana) is the largest lake in Ethiopia and the source of the Blue Nile. Located in Amhara Region in the north-western Ethiopian Highlands, the lake is approximately long and wid ...
in Ethiopia. It has also been possibly recorded in southeastern Sudan and in extreme northwestern Kenya. It has taken over the habitat that was occupied by the Abyssinian hare. It inhabits grasslands,
steppe In physical geography, a steppe () is an ecoregion characterized by grassland plains without trees apart from those near rivers and lakes. Steppe biomes may include: * the montane grasslands and shrublands biome * the temperate grasslands, ...
s, grassy parts of forests, and forest peripheries. It prefers a relatively less open habitat than that of the Abyssinian hare, and similar to the preference by the African savanna hare in Kenya. It is found at elevations of above sea level. Its distribution is
allopatric Allopatric speciation () – also referred to as geographic speciation, vicariant speciation, or its earlier name the dumbbell model – is a mode of speciation that occurs when biological populations become geographically isolated from ...
or
parapatric In parapatric speciation, two subpopulations of a species evolve reproductive isolation from one another while continuing to exchange genes. This mode of speciation has three distinguishing characteristics: 1) mating occurs non-randomly, 2) gene ...
with that of the African savanna hare.


Behaviour and ecology

No data regarding its behaviour, ecology, or reproduction has been recorded. The spotted hyena (''
Crocuta crocuta The spotted hyena (''Crocuta crocuta''), also known as the laughing hyena, is a hyena species, currently classed as the sole extant member of the genus ''Crocuta'', native to sub-Saharan Africa. It is listed as being of least concern by the IUC ...
'') is a predator of the Ethiopian hare. The tick ''
Rhipicephalus pulchellus The zebra tick or yellow back tick (''Rhipicephalus pulchellus'') is a species of hard tick. It is common in the Horn of Africa, with a habitat of the Rift Valley and eastward. It feeds upon a wide variety of species, including livestock, wild m ...
'' has been observed
parasitizing Parasitism is a Symbiosis, close relationship between species, where one organism, the parasite, lives on or inside another organism, the Host (biology), host, causing it some harm, and is Adaptation, adapted structurally to this way of lif ...
the Ethiopian hare.


Status and conservation

Since 1996, the status of the Ethiopian hare has been 
data deficient A data deficient (DD) species is one which has been categorized by the International Union for Conservation of Nature (IUCN) as offering insufficient information for a proper assessment of conservation status to be made. This does not necessaril ...
on the 
IUCN Red List of Endangered Species The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biolog ...
. This is because even though it is widespread species, very little is known about its status and ecology. The current state of its population trend is unclear. There are no known threats to the species. It occurs in the
Abijatta-Shalla National Park Abijatta-Shalla National Park is a national park in Ethiopia. It is located in the Oromia Region and the Ethiopian Highlands region, 200 kilometers south of Addis Ababa, and east of the Batu–Shashamane highway. History Although its intent w ...
and the
Gambela National Park Gambella National Park, also spelled Gambela National Park, is a large national park in Ethiopia. It is the nation's largest national park and is located several hundred kilometers from Addis Ababa. It was established in 1974, but is not fully pr ...
.


Notes


References

{{Taxonbar, from=Q257743 Lepus Mammals described in 1903 Taxa named by Oldfield Thomas Mammals of Ethiopia Taxonomy articles created by Polbot Endemic fauna of Ethiopia