''Etheostoma variatum'', the variegate darter, is a species of freshwater

ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or hor ...

, a darter from the

subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...

Etheostomatinae Etheosomatidae is a species rich subfamily of freshwater ray-finned fish, the members of which are commonly known as the darters. The subfamily is part of the family Percidae which also includes the perches, ruffes and pikeperches. The family i ...

, part of the

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Percidae The Percidae are a family of ray-finned fish, part of the order Perciformes, which are found in fresh and brackish waters of the Northern Hemisphere. The majority are Nearctic, but there are also Palearctic species. The family contains more than ...

, which also contains the

perch Perch is a common name for fish of the genus ''Perca'', freshwater gamefish belonging to the family Percidae. The perch, of which three species occur in different geographical areas, lend their name to a large order of vertebrates: the Percif ...

es, ruffes and pikeperches. It is native to the eastern United States where it occurs primarily in the

Ohio River The Ohio River is a long river in the United States. It is located at the boundary of the Midwestern and Southern United States, flowing southwesterly from western Pennsylvania to its mouth on the Mississippi River at the southern tip of Illino ...

basin; it is present in the states of

Ohio Ohio () is a state in the Midwestern region of the United States. Of the fifty U.S. states, it is the 34th-largest by area, and with a population of nearly 11.8 million, is the seventh-most populous and tenth-most densely populated. The sta ...

New York New York most commonly refers to: * New York City, the most populous city in the United States, located in the state of New York * New York (state), a state in the northeastern United States New York may also refer to: Film and television * '' ...

Pennsylvania Pennsylvania (; ( Pennsylvania Dutch: )), officially the Commonwealth of Pennsylvania, is a state spanning the Mid-Atlantic, Northeastern, Appalachian, and Great Lakes regions of the United States. It borders Delaware to its southeast, ...

Indiana Indiana () is a U.S. state in the Midwestern United States. It is the 38th-largest by area and the 17th-most populous of the 50 States. Its capital and largest city is Indianapolis. Indiana was admitted to the United States as the 19th s ...

West Virginia West Virginia is a state in the Appalachian, Mid-Atlantic and Southeastern regions of the United States.The Census Bureau and the Association of American Geographers classify West Virginia as part of the Southern United States while the Bur ...

and

Kentucky Kentucky ( , ), officially the Commonwealth of Kentucky, is a state in the Southeastern region of the United States and one of the states of the Upper South. It borders Illinois, Indiana, and Ohio to the north; West Virginia and Virginia to ...

with a small population in

Virginia Virginia, officially the Commonwealth of Virginia, is a state in the Mid-Atlantic and Southeastern regions of the United States, between the Atlantic Coast and the Appalachian Mountains. The geography and climate of the Commonwealth ar ...

. It is a large darter, growing to long, orange-red with black vertical barring on its body, and horizontal barring on its front dorsal fin. Its typical habitat is swift-flowing riffles with rubble, boulder and gravel. No major threats have been identified for this fish and the

International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...

has assessed its conservation status as being of "

least concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. T ...

". Biology

Description

''Etheostoma variatum'' is large in size for its family; it has an average length of between . It has a wide variety of markings. There is a red-orange band on its dorsal fin, which is followed by a blue-green band and a red-brown band at the edge of the fin. There are also red spots located on its other fins. This darter has large pectoral fins, which are used to keep the fish upright in the fast-moving streams it inhabits. It also has four to six saddles along its back. Three to four of these saddles tend to be dark and visible while the remaining saddles are not.

Habitat

''Etheostoma variatum'' lives in small and medium rivers and streams. They tend to live within

riffles A riffle is a shallow landform in a flowing channel. Colloquially, it is a shallow place in a river where water flows quickly past rocks. However, in geology a riffle has specific characteristics. Topographic, sedimentary and hydraulic indica ...

that contain small rocks. The darter will tend to live in the fastest flowing riffles. In order to survive, it needs rivers and streams with high water quality. Due to this attribute, the fish is often viewed as an indicator of good water quality. The darter lives predominantly in the states of

and

. It has also been found within

, although only rarely.

Behavior

Diet

The diet of ''E. variatum'' consists of

crustaceans Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...

, insect larvae and other aquatic

invertebrates Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate ...

Reproduction

During the early spring, males establish spawning territory, which will occur from April to May. After spawning, the eggs are laid in riffles by females, who will bury them with pebbles. After the eggs are buried, they are abandoned by the parents.

Status

The population trend of this fish seems to be stable and it is a common species with numerous sub-populations over a wide range, and the

has assessed its conservation status as being of "

References

{{Taxonbar, from=Q3761058 variatum Fish described in 1819 Freshwater fish of the United States Endemic fauna of the United States