Etheostoma Coosae
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The Coosa darter (''Etheostoma coosae'') is a species of freshwater ray-finned fish, a darter from the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
Etheostomatinae Etheosomatidae is a species rich subfamily of freshwater ray-finned fish, the members of which are commonly known as the darters. The subfamily is part of the family Percidae which also includes the perches, ruffes and pikeperches. The famil ...
, part of the family Percidae, which also contains the perches, ruffes and pikeperches. It is endemic to the eastern United States.


Description

The Coosa darter is a robust species of darter which can be identified by having a blunt snout with a small mouth. The spiny part of its dorsal fin is marked with bands of color and has a central red band along the whole of its length, although it does not possess the anterior ocellus found in many other species in the
subgenus In biology, a subgenus (plural: subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between t ...
''Ulocentra''. Above and below this central band there are alternating clear and dark bans. In the soft-rayed part of the dorsal fin, the middle part of the membranes between each ray is red. The color of the body is yellow-olive, marked with 8-9 dark blotches located dorsally and on the flanks. The lower snout and the throat are pale green, while the
anal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
and upper and lower lobes of the caudal fin are turquoise. The blotches along the flank have a slightly green hue. The maximum recorded total length for this species is , although the
standard length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish m ...
of males is more commonly around and of females .


Distribution

The Coosa darter is only found in the Coosa River system, which drains into Mobile Bay in Georgia, Alabama and Tennessee.


Habitat and biology

The Coosa darter is found in rocky pools and nearby riffles of creeks and small to medium rivers, as well as in streams. It feeds mainly on the larvae of
midge A midge is any small fly, including species in several families of non-mosquito Nematoceran Diptera. Midges are found (seasonally or otherwise) on practically every land area outside permanently arid deserts and the frigid zones. Some mid ...
and blackfly larvae, with smaller amounts of cladocera,
copepods Copepods (; meaning "oar-feet") are a group of small crustaceans found in nearly every freshwater and saltwater habitat. Some species are planktonic (inhabiting sea waters), some are benthic (living on the ocean floor), a number of species have p ...
,
mayfly Mayflies (also known as shadflies or fishflies in Canada and the upper Midwestern United States, as Canadian soldiers in the American Great Lakes region, and as up-winged flies in the United Kingdom) are aquatic insects belonging to the ord ...
nymphs, and caddisfly larvae. Insects, especially flies, are more important in summer and crustaceans become more important in the winter. The Coosa darter is known to spawn from mid-March to mid-May. The female lays a single egg at a time, placing it in small cracks and crevices in wood, rocks, or other hard substrates. The male fertilizes the eggs as soon as they are laid. They may spawn in any position from horizontal to vertical.


Taxonomy

The Coosa darter was first formally described as ''Poecilichthys coosae'' in 1945 by the
American American(s) may refer to: * American, something of, from, or related to the United States of America, commonly known as the "United States" or "America" ** Americans, citizens and nationals of the United States of America ** American ancestry, pe ...
ichthyologist Ichthyology is the branch of zoology devoted to the study of fish, including bony fish ( Osteichthyes), cartilaginous fish (Chondrichthyes), and jawless fish (Agnatha). According to FishBase, 33,400 species of fish had been described as of Octobe ...
Henry Weed Fowler (1878-1965) with the
type locality Type locality may refer to: * Type locality (biology) * Type locality (geology) See also * Local (disambiguation) * Locality (disambiguation) {{disambiguation ...
given as a small creek in the Coosa River drainage within Cherokee County, Alabama, near the settlement of Chesterfield.


Status

The
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
has listed this species as being of " Least Concern" because it has an extensive range in the Coosa River system, has a large total population size, and has numerous subpopulations. In general, the population trend seems stable and no major threats have been identified.


References

*O'Neal, P. "Life history of the ''Etheostoma-coosae'' ''Pisces Percidae'' in Barbaree Creek Alabama U.S.." Tulane Studies in Zoology and Botany. 23.1 (1981): 75–84. Web. 23 Sep. 2013. *https://web.archive.org/web/20111002015239/http://www.bio.utk.edu/hulseylab/Fishlist.html *Ryon, Michael G. "The life history and ecology of ''Etheostoma trisella'' (Pisces: Percidae)." American Midland Naturalist (1986): 73–86. *Brogdon, Stephen M., Christopher R. Tabit, and Leos G. Kral. "Population structure of the Tallapoosa darter (''Etheostoma tallapoosae'')." Southeastern Naturalist 2.4 (2003): 487–498. *Bart, Henry L. "Spawning behavior of ''Etheostoma davisoni'' Hay." Copeia 1992.2 (1992): 537–539. *Storey, Casey Michael. Genetic population structure and life history aspects of the federally threatened Cherokee darter, ''Etheostoma scotti''. Diss. uga, 2003. {{Taxonbar, from=Q3753306 Etheostoma Fish described in 1945