''Etelis carbunculus'', the deep-water red snapper, ruby snapper or longtail snapper, is a species of

ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or hor ...

, a snapper belonging to the

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Lutjanidae Lutjanidae, or snappers are a family of perciform fish, mainly marine, but with some members inhabiting estuaries, feeding in fresh water. The family includes about 113 species. Some are important food fish. One of the best known is the red snapp ...

. It is found in the

Indo-Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...

region.

Description

''Etelis carbunculus'' is an elongated fish with a small head and a large eye, the space between the eyes is flat. The mouth extends back as far as the middle of the eye and the jaws are each equipped with a single row of conical teeth with 1 or 2 pairs of enlarged canines at the front. The

vomerine teeth The vomer (; lat, vomer, lit=ploughshare) is one of the unpaired facial bones of the skull. It is located in the midsagittal line, and articulates with the sphenoid, the ethmoid, the left and right palatine bones, and the left and right maxill ...

are arranged in a slender V-shaped patch. The forked

caudal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...

has relatively short lobes in comparison to congeners. It has a continuous

dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through conv ...

which has a deep notch at the junction of its spiny part and the ultimate soft ray of both the dorsal and

anal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...

s extends beyond the membrane, being longer than the penultimate ray. The dorsal fin contains 10 spines and 11 soft rays while the anal fin has 3 spines and 8 soft rays, both fins lacking scales. The

pectoral fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as ...

s are long, almost equal to the head length. The maximum recorded

fork length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish m ...

is , although a

total length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish ...

of is more typical. The overall colour of this species is reddish to pinkish with a whitish abdomen. There are red margins to the dorsal fin and the caudal fin which has a white tip to its lower lobe.

Distribution

''Etelis carbunculus'' has a wide distribution and can be found throughout the Indian and Pacific Oceans. It occurs from the

Red Sea The Red Sea ( ar, البحر الأحمر - بحر القلزم, translit=Modern: al-Baḥr al-ʾAḥmar, Medieval: Baḥr al-Qulzum; or ; Coptic: ⲫⲓⲟⲙ ⲛ̀ϩⲁϩ ''Phiom Enhah'' or ⲫⲓⲟⲙ ⲛ̀ϣⲁⲣⲓ ''Phiom ǹšari''; T ...

south to Mozambique on the coast of East Africa to the

Persian Gulf The Persian Gulf ( fa, خلیج فارس, translit=xalij-e fârs, lit=Gulf of Persis, Fars, ), sometimes called the ( ar, اَلْخَلِيْجُ ٱلْعَرَبِيُّ, Al-Khalīj al-ˁArabī), is a Mediterranean sea (oceanography), me ...

, across the Indian Ocean into the Pacific Ocean. In the Pacific its range extends north to Japan, south to Australia and east to

Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only stat ...

. It has also been found around Three Kings Island off northern New Zealand. In Australia it is found from

Geraldton Geraldton (Wajarri: ''Jambinu'', Wilunyu: ''Jambinbirri'') is a coastal city in the Mid West region of the Australian state of Western Australia, north of the state capital, Perth. At June 2018, Geraldton had an urban population of 37,648. ...

Western Australia Western Australia (commonly abbreviated as WA) is a state of Australia occupying the western percent of the land area of Australia excluding external territories. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to th ...

east into the

Arafura Sea The Arafura Sea (or Arafuru Sea) lies west of the Pacific Ocean, overlying the continental shelf between Australia and Western New Guinea (also called Papua), which is the Indonesian part of the Island of New Guinea. Geography The Arafura Sea is ...

to the northeast of Darwin in the

Northern Territory The Northern Territory (commonly abbreviated as NT; formally the Northern Territory of Australia) is an states and territories of Australia, Australian territory in the central and central northern regions of Australia. The Northern Territory ...

onto the outer

Great Barrier Reef The Great Barrier Reef is the world's largest coral reef system composed of over 2,900 individual reefs and 900 islands stretching for over over an area of approximately . The reef is located in the Coral Sea, off the coast of Queensland, ...

off

Queensland ) , nickname = Sunshine State , image_map = Queensland in Australia.svg , map_caption = Location of Queensland in Australia , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , established_ ...

, maybe also occurring farther south. It also occurs at

Lord Howe Island Lord Howe Island (; formerly Lord Howe's Island) is an irregularly crescent-shaped volcanic remnant in the Tasman Sea between Australia and New Zealand, part of the Australian state of New South Wales. It lies directly east of mainland P ...

in the

Tasman Sea The Tasman Sea (Māori: ''Te Tai-o-Rēhua'', ) is a marginal sea of the South Pacific Ocean, situated between Australia and New Zealand. It measures about across and about from north to south. The sea was named after the Dutch explorer Abe ...

as well as at

Christmas Island Christmas Island, officially the Territory of Christmas Island, is an Australian external territory comprising the island of the same name. It is located in the Indian Ocean, around south of Java and Sumatra and around north-west of the ...

and

Cocos (Keeling) Island ) , anthem = "''Advance Australia Fair''" , song_type = , song = , image_map = Australia on the globe (Cocos (Keeling) Islands special) (Southeast Asia centered).svg , map_alt = Location of the Cocos (Keeling) Islands , map_caption = ...

Habitat and biology

''Etelis carbunculus'' is found over rock substrates and rocky reefs at depths between , in or around the benthos. It has been reported forming aggregations. It is a predatory species feeding on fishes,

squid True squid are molluscs with an elongated soft body, large eyes, eight arms, and two tentacles in the superorder Decapodiformes, though many other molluscs within the broader Neocoleoidea are also called squid despite not strictly fitting t ...

crustaceans Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...

and

zooplankton Zooplankton are the animal component of the planktonic community ("zoo" comes from the Greek word for ''animal''). Plankton are aquatic organisms that are unable to swim effectively against currents, and consequently drift or are carried along by ...

. They breed all year round in Vanuatu,

spawning Spawn is the eggs and sperm released or deposited into water by aquatic animals. As a verb, ''to spawn'' refers to the process of releasing the eggs and sperm, and the act of both sexes is called spawning. Most aquatic animals, except for aquati ...

peaking there in November.

Parasites

The

monopisthocotylea The Monopisthocotylea are a subclass of parasitic flatworms in the class Monogenea. WoRMS (2019). Monopisthocotylea. Accessed at: http://www.marinespecies.org/aphia.php?p=taxdetails&id=119219 on 2019-02-08 Yamaguti, S. (1963). Systema Helminthum V ...

monogenea Monogeneans are a group of ectoparasitic flatworms commonly found on the skin, gills, or fins of fish. They have a direct lifecycle and do not require an intermediate host. Adults are hermaphrodites, meaning they have both male and female reprod ...

n of the genus ''

Lagenivaginopseudobenedenia ''Lagenivaginopseudobenedenia'' is a genus of monopisthocotylean monogenean (a group of ectoparasitic flatworms), included in the family Capsalidae.Whittington, I. D. 2004: The Capsalidae (Monogenea: Monopisthocotylea): a review of diversity, cla ...

'' are known to

parasitise Parasitism is a close relationship between species, where one organism, the parasite, lives on or inside another organism, the host, causing it some harm, and is adapted structurally to this way of life. The entomologist E. O. Wilson has ...

''E. carbunculus''. These parasites have what is possibly the longest valid generic name of the

zoological nomenclature The International Code of Zoological Nomenclature (ICZN) is a widely accepted convention in zoology that rules the formal scientific naming of organisms treated as animals. It is also informally known as the ICZN Code, for its publisher, the In ...

for a non-fossil organism.

Taxonomy

''Etelis carbunculus'' was first formally described in 1828 by the

French French (french: français(e), link=no) may refer to: * Something of, from, or related to France ** French language, which originated in France, and its various dialects and accents ** French people, a nation and ethnic group identified with Franc ...

zoologist Zoology ()The pronunciation of zoology as is usually regarded as nonstandard, though it is not uncommon. is the branch of biology that studies the Animal, animal kingdom, including the anatomy, structure, embryology, evolution, Biological clas ...

Georges Cuvier with the type locality given as Mahé in the Seychelles. At the time it was the only species in the

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

'' Etelis'' so it is the

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

of that genus. The

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

''carbunculus'' is a “ruby red precious stone”, a reference to the reddish colour of this fish.

Fisheries

''Etelis carbunculus'' is a target for fisheries throughout its range and the stocks have been reported to be declining in some regions. It is caught using bottom longlines and deep handlines and is an important food fish. In Australia its stock is managed under the Western Deep Water Fishery. The caught fish are sold fresh or frozen.

References

{{taxonbar, from=Q1042554 carbunculus Fish described in 1828