Eschrichtioides Portis 1885 Fig 2
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''Eschrichtioides'' is an extinct
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
baleen whale Baleen whales (systematic name Mysticeti), also known as whalebone whales, are a parvorder of carnivorous marine mammals of the infraorder Cetacea (whales, dolphins and porpoises) which use keratinaceous baleen plates (or "whalebone") in their ...
known from the early
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...
. Its type species, ''E. gastaldii'', had a complex taxonomic history, starting as a cetothere, then as an extinct member of ''
Balaenoptera ''Balaenoptera'' () is a genus of rorquals containing eight extant species. ''Balaenoptera'' comprises all but two of the extant species in its family (the humpback whale and gray whale); the genus is currently polyphyletic, with the two aforem ...
'', before being finally recognized as a relative of the
gray whale The gray whale (''Eschrichtius robustus''), also known as the grey whale,Britannica Micro.: v. IV, p. 693. gray back whale, Pacific gray whale, Korean gray whale, or California gray whale, is a baleen whale that migrates between feeding and bree ...
.


Distribution

''Eschrichtioides'' is one of two '' Eschrichtius'' relatives known from the
Neogene The Neogene ( ), informally Upper Tertiary or Late Tertiary, is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period million years ago ( Mya) to the beginning of the present Quaternary Period Mya. ...
of Italy, the other being '' Archaeschrichtius''. Its
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
, MRSN 13802, comes from the early Pliocene-age Sabbie d'Asti Formation of the
Piedmont it, Piemontese , population_note = , population_blank1_title = , population_blank1 = , demographics_type1 = , demographics1_footnotes = , demographics1_title1 = , demographics1_info1 = , demographics1_title2 ...
region in Italy.


Classification

Like other cetaceans from the Italian Pliocene, the classification of ''Eschrichtioides'' was highly contentious. described several species of ''
Cetotherium ''Cetotherium'' ("whale beast") is an extinct genus of baleen whales from the family Cetotheriidae. Taxonomy The family Cetotheriidae and the genus ''Cetotherium'' (sensu lato) have been used as wastebaskets for all kinds of baleen whales, mo ...
'' from the Pliocene of northern Italy, including ''C. cortesi'' based on a specimen (
MSRN {{Unreferenced, date=August 2009 The Mobile Station Roaming Number (MSRN) is an E.164 E.164 is an international standard (ITU-T Recommendation), titled ''The international public telecommunication numbering plan'', that defines a numbering plan ...
 13802) found by Italian geologist
Giuseppe Cortesi Giuseppe is the Italian form of the given name Joseph, from Latin Iōsēphus from Ancient Greek Ἰωσήφ (Iōsḗph), from Hebrew יוסף. It is the most common name in Italy and is unique (97%) to it. The feminine form of the name is Giuse ...
in 1816 near the town
Cortandone Cortandone is a ''comune'' (municipality) in the Province of Asti in the Italian region Piedmont, located about southeast of Turin and about northwest of Asti. As of 31 December 2016, it had a population of 331 and an area of .All demographics an ...
(: paleocoordinates ) described how he, after a day of searching for fossils, found what he first thought was a small piece of wood in a small river. After a closer inspection it proved to be a bone, and Cortesi and his companion found more and more of them in the banks of the river. After four days of digging they had unveiled a giant skeleton easily identifiable as a whale 12 feet 5 inches long. The three feet long head and its dentaries had no signs of either teeth nor
tooth sockets Dental alveoli (singular ''alveolus'') are sockets in the jaws in which the roots of teeth are held in the alveolar process with the periodontal ligament. The lay term for dental alveoli is tooth sockets. A joint that connects the roots of the ...
. Cortesi noted that at that time few naturalists could assign cetacean fossils to individual species, and he therefore never named his specimen. thought the "Baleine de Cortesi" represented a distinctive species because it was a very small adult individual and because the curvature of the maxillary branches was less convex than in any other known whale. Other French naturalists named it "Le rorqual de Cortési"; arguing that both Cortesi's fossil whales ("rorqual de Cortési" and "de Cuvier") were closely related to extant rorquals and the only difference between them was the much smaller size of the former. assigned the "baleine de Cortesi" to his own genus ''
Cetotherium ''Cetotherium'' ("whale beast") is an extinct genus of baleen whales from the family Cetotheriidae. Taxonomy The family Cetotheriidae and the genus ''Cetotherium'' (sensu lato) have been used as wastebaskets for all kinds of baleen whales, mo ...
Cortesii''. In a preliminary note pointed out differences in cranial morphology in two skulls both described as "''Cetotherium cortesii''" (a species named and described by ), and he therefore proposed the new specific name "''Cetotherium gastaldii''" (in honour of Italian palaeontologist
Bartolomeo Gastaldi Bartolomeo Gastaldi (10 February 1818 – 5 January 1879) was an Italian geologist and palæontologist, and one of the founders of the Club Alpino Italiano. Gastaldi was born in Turin, then capital of the Kingdom of Sardinia. As a child he develo ...
) for one of the specimens. described the specimen and reiterated his arguments from 1875. Originally misidentified as the balaenopterid "Balaena" ''cortesii'', described "B." ''cortesii'' as a species of ''Cetotherium''. finally named it ''C. gastaldii''. However, several authors considered it to be referrable to ''
Balaenoptera ''Balaenoptera'' () is a genus of rorquals containing eight extant species. ''Balaenoptera'' comprises all but two of the extant species in its family (the humpback whale and gray whale); the genus is currently polyphyletic, with the two aforem ...
'', a taxonomic arrangement which was accepted for more than a century. However, re-examination of MRSN 13802 revealed that "C." gastaldii was not a balaenopterid and instead constituted its own genus within
Eschrichtiidae Eschrichtiidae or the gray whales is a family of baleen whale (Parvorder Mysticeti) with a single extant species, the gray whale (''Eschrichtius robustus''), as well as three described fossil genera: '' Archaeschrichtius'' and ''Eschrichtioide ...
.


References


Notes


Sources

* ** * ** * * ** * * * ** * * ** * ** *
Plates 1, 2, 5
** * {{Taxonbar, from=Q15701238 Pliocene mammals of Europe Pliocene cetaceans Fossil taxa described in 2008 Baleen whales Prehistoric cetacean genera