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''Erica'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
of roughly 857
species In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...
of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. They include all forbs (flowering plants without a woody stem), grasses and grass-like plants, a vast majority of ...
s in the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Ericaceae. The English common names heath and heather are shared by some closely related genera of similar appearance. The genus ''
Calluna ''Calluna vulgaris'', common heather, ling, or simply heather, is the sole species in the genus ''Calluna'' in the flowering plant family Ericaceae. It is a low-growing evergreen shrub growing to tall, or rarely to and taller, and is found wi ...
'' was formerly included in ''Erica'' – it differs in having even smaller scale-leaves (less than 2–3 mm long), and the flower
corolla Corolla may refer to: *Corolla (botany), the petals of a flower, considered as a unit *Toyota Corolla, an automobile model name * Corolla (headgear), an ancient headdress in the form of a circlet or crown * ''Corolla'' (gastropod), a genus of moll ...
consisting of separate petals. ''Erica'' is sometimes referred to as "winter (or spring) heather" to distinguish it from ''Calluna'' "summer (or autumn) heather".


Etymology

The
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power ...
word ''erica'' means "heath" or "broom". It is believed that Pliny adapted ''erica'' from
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic p ...
ἐρείκη. The expected Anglo-Latin pronunciation, , may be given in dictionaries ('' OED'': "Erica"), but is more commonly heard.


Description

Most of the species of ''Erica'' are small shrubs from high, though some are taller; the tallest are '' E. arborea'' (tree heath) and '' E. scoparia'' (besom heath), both of which can reach up to tall. All are
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, which ...
, with minute, needle-like leaves long. Flowers are sometimes axillary, and sometimes borne in terminal umbels or spikes, and are usually outward or downward facing. The
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiosper ...
s are very small, and in some species may survive in the soil for decades.


Taxonomy and phylogeny

Dulfer published the last revision of the genus ''Erica'' in the 1960s, treating 605 species. Many new species have subsequently been described (particularly in South Africa) and a further 83 have been included in ''Erica'' from former “minor genera”, such as ''Phillipia'' Klotzsch and ''Blaeria'' L. A more recent overview of ''Erica'' species is provided in an electronic identification aid, but a modern taxonomic revision of the genus as a whole is still lacking. A number of increasingly detailed phylogenetic hypotheses for ''Erica'' have been published based on nuclear
ribosomal Ribosomes ( ) are macromolecular machines, found within all cells, that perform biological protein synthesis (mRNA translation). Ribosomes link amino acids together in the order specified by the codons of messenger RNA (mRNA) molecules to ...
and
plastid The plastid (Greek: πλαστός; plastós: formed, molded – plural plastids) is a membrane-bound organelle found in the cells of plants, algae, and some other eukaryotic organisms. They are considered to be intracellular endosymbiotic cyanobac ...
DNA sequences. The closest relatives of ''Erica'' are '' Daboecia'' (one or two species) and ''
Calluna ''Calluna vulgaris'', common heather, ling, or simply heather, is the sole species in the genus ''Calluna'' in the flowering plant family Ericaceae. It is a low-growing evergreen shrub growing to tall, or rarely to and taller, and is found wi ...
'' (monospecific), representing the oldest surviving lineages of a, by inference, ancestrally Palearctic tribe Ericeae. The small number of European ''Erica'' species represent the oldest lineages of the genus, within which a single, order-of-magnitude more species-rich, African clade is nested. Within the African clade, Cape and Madagascan/Mascarene species respectively represent
monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic ...
groups.


Species

Selected species include:


Habitat

Around 690 of the species are
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found els ...
to South Africa, and these are often called the Cape heaths, forming the largest genus in the fynbos. The remaining species are native to other parts of Africa, Madagascar, the Mediterranean, and Europe. Like most Ericaceae, ''Erica'' species are mainly calcifuges, being limited to acidic or very acidic soils. In fact, the term "ericaceous" is frequently applied to all calcifuges, and to the
compost Compost is a mixture of ingredients used as plant fertilizer and to improve soil's physical, chemical and biological properties. It is commonly prepared by decomposing plant, food waste, recycling organic materials and manure. The resulting ...
used in their cultivation. Soils range from dry, sandy soils to extremely wet ones such as
bog A bog or bogland is a wetland that accumulates peat as a deposit of dead plant materials often mosses, typically sphagnum moss. It is one of the four main types of wetlands. Other names for bogs include mire, mosses, quagmire, and muskeg; a ...
. They often dominate dwarf-shrub habitats ( heathland and moorland), or the ground vegetation of open acidic
woodland A woodland () is, in the broad sense, land covered with trees, or in a narrow sense, synonymous with wood (or in the U.S., the '' plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunlight and limited shade (see ...
.


Ecology

Plants of this genus are eaten mainly by the
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. Th ...
e of many Lepidoptera species, including emperor moth, garden tiger moth,
true lover's knot The term true lover's knot, also called true love knot, is used for many distinct knots. The association of knots with the symbolism of love, friendship and affection dates back to antiquity. Because of this, no single knot can be labeled the ...
, wormwood pug, the silver-studded blue, and the ''
Coleophora ''Coleophora'' is a very large genus of moths of the family Coleophoridae. It contains some 1,350 described species. The genus is represented on all continents, but the majority are found in the Nearctic and Palaearctic regions. Many authors h ...
'' case-bearers '' C. juncicolella'' and '' C. pyrrhulipennella''. Some species of sunbirds are known to visit and pollinate ''Erica''. Two such species are the southern double-collared sunbird and the orange-breasted sunbird.


Cultivation

''Erica'' species are grown as landscape or garden plants for their floral effect. They associate well with conifers and are frequently seen in planting schemes as massed groundcover beneath varieties of dwarf conifers. They are capable of producing flower colour throughout the year. They can also be grown in tubs or window boxes to provide interest through autumn and into winter.


References

{{Taxonbar, from=Q206998 Ericaceae genera Groundcovers Taxa named by Carl Linnaeus Subshrubs