Eremocarya Lepida
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''Eremocarya'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
s in the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Boraginaceae Boraginaceae, the borage or forget-me-not family, includes about 2,000 species of shrubs, trees and herbs in 146, to 156 genera with a worldwide distribution. The APG IV system from 2016 classifies the Boraginaceae as single family of the order ...
. There are about 63 species and its native range extends through western United States to northwestern Mexico. It is part of subtribe of ''Amsinckiinae''. It was once thought to be an either a subgenus or synonym of ''
Cryptantha ''Cryptantha'' is a genus of flowering plants in the borage family, Boraginaceae. They are known commonly as cat's eyes and popcorn flowers (the latter name is also used to refer to the closely related genus ''Plagiobothrys'',Hasenstab-Lehman, ...
'' , before being segregated out due to molecular phylogenetic analysis.


Description

A profusely branching annual herb with very slender, ascending, nearly leafless stems and the leaves are arranged in a basal rosette. The roots and the lower parts of the stems are often stained with a red, or purple hue. The flowers in March–June, are dense
racemes A raceme ( or ) or racemoid is an unbranched, indeterminate type of inflorescence bearing flowers having short floral stalks along the shoots that bear the flowers. The oldest flowers grow close to the base and new flowers are produced as the s ...
, spiciform (spike-shaped) with evenly spaced, leafy-bracteate beneath each flower. The
calyx Calyx or calyce (plural "calyces"), from the Latin ''calix'' which itself comes from the Ancient Greek ''κάλυξ'' (''kálux'') meaning "husk" or "pod", may refer to: Biology * Calyx (anatomy), collective name for several cup-like structures ...
is small and divided into 5 sections from the base. It has a small white
corolla Corolla may refer to: *Corolla (botany), the petals of a flower, considered as a unit *Toyota Corolla, an automobile model name * Corolla (headgear), an ancient headdress in the form of a circlet or crown * ''Corolla'' (gastropod), a genus of moll ...
. It has 4
ovules In seed plants, the ovule is the structure that gives rise to and contains the female reproductive cells. It consists of three parts: the ''integument'', forming its outer layer, the ''nucellus'' (or remnant of the megasporangium), and the fe ...
and 4 nutlets (which appear after flowering), which are similar in size and shape. The gynobase (a short conical or flat elevation of the receptacle of a flower, bearing the
gynoecium Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) ''pistils'' ...
) is thin and columnar, they are nearly similar (in form) to the wide style, which is dilated and wider that the stigma when in fruit.Forrest Shreve and Ira Loren Wiggins


Taxonomy

The Latin
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''Eremocarya'' is derived from "Eremos" which is Greek for "desert" or "lonely" and "caryum" is Greek for "nut". It was first published by
Edward Lee Greene Edward Lee Greene (August 20, 1843–November 10, 1915) was an American botanist known for his numerous publications including the two-part ''Landmarks of Botanical History'' and the describing of over 4,400 species of plants in the American W ...
in Pittonia vol.1 on page 58 in 1887. Then in 1924, Ivan M. Johnston wrote that the genus of ''Oreocarya'' could be combined with ''Cryptantha''. Edwin Blake Payson in 1927 (A Monograph of the section ''Oreocarya'' of ''Cryptantha'', Ann. Mo. Bot. Gard. 14:211-358) agreed with Johnston and he had four sections in ''Cryptantha'': ''Eucryptantha'' (= ''Cryptantha''), ''Geocarya'', ''Krynitzkia'' (inclusive of ''Eremocarya'', '' Greeneocharis'', and '' Johnstonella''), and ''Oreocarya''. Larry Higgins (1971), another expert on the perennial taxa, published a revised monograph of ''Oreocarya'', and agreed with Johnston and Payson on the inclusion of ''Oreocarya'' within ''Cryptantha'', but also elevating the four sections of Johnston (1927) and Payson (1927) to
subgenera In biology, a subgenus (plural: subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between t ...
. Although they were sometimes still called synonyms of ''Cryptantha''. In 2012, the
phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
relationship of members of the genus ''Cryptantha'' was carried out, based on
dna sequencing DNA sequencing is the process of determining the nucleic acid sequence – the order of nucleotides in DNA. It includes any method or technology that is used to determine the order of the four bases: adenine, guanine, cytosine, and thymine. Th ...
analyses, it was then proposed that the resurrection of the following genera ''Eremocarya'', ''Greeneocharis'', ''Johnstonella'', and also '' Oreocarya''.


Species

2 accepted species; * '' Eremocarya lepida'' * '' Eremocarya micrantha'' In 2016, a large flowered version of ''Eremocarya micrantha'' was found and published as ''E. micrantha'' var. ''pseudolepida''.


Distribution

The genus is found in the United States (within the states of
Arizona Arizona ( ; nv, Hoozdo Hahoodzo ; ood, Alĭ ṣonak ) is a state in the Southwestern United States. It is the 6th largest and the 14th most populous of the 50 states. Its capital and largest city is Phoenix. Arizona is part of the Fou ...
,
California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...
,
Nevada Nevada ( ; ) is a U.S. state, state in the Western United States, Western region of the United States. It is bordered by Oregon to the northwest, Idaho to the northeast, California to the west, Arizona to the southeast, and Utah to the east. N ...
,
New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe , LargestCity = Albuquerque , LargestMetro = Tiguex , OfficialLang = None , Languages = English, Spanish ( New Mexican), Navajo, Ker ...
,
Oregon Oregon () is a U.S. state, state in the Pacific Northwest region of the Western United States. The Columbia River delineates much of Oregon's northern boundary with Washington (state), Washington, while the Snake River delineates much of it ...
,
Texas Texas (, ; Spanish language, Spanish: ''Texas'', ''Tejas'') is a state in the South Central United States, South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2 ...
and
Utah Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to it ...
) and also in north western
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
.


Habitat

Species of the genus are found in dry sandy places at altitudes of below above sea level or on montane slopes and valley sides at .Philip Alexander Munz (and David D.Keck)


References


Other sources

* Cronquist, A. et al. 1972-. Intermountain flora. * Johnston, I. M. 1927. Studies in the Boraginaceae VI. A revision of the South American Boraginoideae. Contr. Gray Herb. 78:31. {{Taxonbar, from=Q21293632 Boraginoideae Boraginaceae genera Plants described in 1887 Flora of the Northwestern United States Flora of the Southwestern United States Flora of the South-Central United States Flora of Northwestern Mexico