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An erection (clinically: penile erection or penile tumescence) is a
physiological Physiology (; ) is the scientific study of functions and mechanisms in a living system. As a sub-discipline of biology, physiology focuses on how organisms, organ systems, individual organs, cells, and biomolecules carry out the chemical ...
phenomenon in which the penis becomes firm, engorged, and enlarged. Penile erection is the result of a complex interaction of psychological, neural, vascular, and endocrine factors, and is often associated with
sexual arousal Sexual arousal (also known as sexual excitement) describes the physiological and psychological responses in preparation for sexual intercourse or when exposed to sexual stimuli. A number of physiological responses occur in the body and mind as ...
or sexual attraction, although erections can also be spontaneous. The shape, angle, and direction of an erection varies considerably between humans. Physiologically, an erection is required for a male to effect vaginal penetration or
sexual intercourse Sexual intercourse (or coitus or copulation) is a sexual activity typically involving the insertion and thrusting of the penis into the vagina for sexual pleasure or reproduction.Sexual intercourse most commonly means penile–vaginal penetrat ...
and is triggered by the
parasympathetic The parasympathetic nervous system (PSNS) is one of the three divisions of the autonomic nervous system, the others being the sympathetic nervous system and the enteric nervous system. The enteric nervous system is sometimes considered part of t ...
division of the
autonomic nervous system The autonomic nervous system (ANS), formerly referred to as the vegetative nervous system, is a division of the peripheral nervous system that supplies viscera, internal organs, smooth muscle and glands. The autonomic nervous system is a control ...
, causing the levels of
nitric oxide Nitric oxide (nitrogen oxide or nitrogen monoxide) is a colorless gas with the formula . It is one of the principal oxides of nitrogen. Nitric oxide is a free radical: it has an unpaired electron, which is sometimes denoted by a dot in its che ...
(a
vasodilator Vasodilation is the widening of blood vessels. It results from relaxation of smooth muscle cells within the vessel walls, in particular in the large veins, large arteries, and smaller arterioles. The process is the opposite of vasoconstriction, ...
) to rise in the
trabecular A trabecula (plural trabeculae, from Latin for "small beam") is a small, often microscopic, tissue element in the form of a small beam, strut or rod that supports or anchors a framework of parts within a body or organ. A trabecula generally has ...
arteries and
smooth muscle Smooth muscle is an involuntary non-striated muscle, so-called because it has no sarcomeres and therefore no striations (''bands'' or ''stripes''). It is divided into two subgroups, single-unit and multiunit smooth muscle. Within single-unit mus ...
of the penis. The arteries dilate causing the corpora cavernosa of the penis (and to a lesser extent the corpus spongiosum) to fill with blood; simultaneously the ischiocavernosus and
bulbospongiosus The bulbospongiosus muscle (bulbocavernosus in older texts) is one of the superficial muscles of the perineum. It has a slightly different origin, insertion and function in males and females. In males, it covers the bulb of the penis. In femal ...
muscles compress the veins of the corpora cavernosa restricting the egress and circulation of this blood. Erection subsides when parasympathetic activity reduces to baseline. As an autonomic nervous system response, an erection may result from a variety of stimuli, including sexual stimulation and
sexual arousal Sexual arousal (also known as sexual excitement) describes the physiological and psychological responses in preparation for sexual intercourse or when exposed to sexual stimuli. A number of physiological responses occur in the body and mind as ...
, and is therefore not entirely under conscious control. Erections during sleep or upon waking up are known as nocturnal penile tumescence (NPT), also known as "morning wood". Absence of nocturnal erection is commonly used to distinguish between physical and psychological causes of
erectile dysfunction Erectile dysfunction (ED), also called impotence, is the type of sexual dysfunction in which the penis fails to become or stay erect during sexual activity. It is the most common sexual problem in men.Cunningham GR, Rosen RC. Overview of male ...
and impotence. The state of a penis which is partly, but not fully, erect is sometimes known as semi-erection (clinically: partial tumescence); a penis which is not erect is typically referred to as being flaccid, or soft.


Physiology

An erection is necessary for
natural insemination Insemination is the introduction of sperm into a female’s reproductive system for the purpose of impregnating, also called fertilizing, the female for sexual reproduction. The sperm is introduced into the uterus of a mammal or the oviduct of an ...
as well as for the harvesting of sperm for artificial insemination, are common for children and infants, and even occur before birth. After reaching puberty, erections occur much more frequently. it occurs when two tubular structures, called the corpora cavernosa, that run the length of the penis, become engorged with venous blood. This may result from any of various
physiological Physiology (; ) is the scientific study of functions and mechanisms in a living system. As a sub-discipline of biology, physiology focuses on how organisms, organ systems, individual organs, cells, and biomolecules carry out the chemical ...
stimuli, also known as sexual stimulation and
sexual arousal Sexual arousal (also known as sexual excitement) describes the physiological and psychological responses in preparation for sexual intercourse or when exposed to sexual stimuli. A number of physiological responses occur in the body and mind as ...
. The corpus spongiosum is a single tubular structure located just below the corpora cavernosa, which contains the urethra, through which urine and semen pass during urination and ejaculation respectively. This may also become slightly engorged with blood, but less so than the corpora cavernosa. The scrotum may, but does not always, become tightened during erection. Generally, in uncircumcised males, the foreskin automatically and gradually retracts, exposing the glans, though some people may have to manually retract their foreskin.


Autonomic control

In the presence of mechanical stimulation, erection is initiated by the
parasympathetic The parasympathetic nervous system (PSNS) is one of the three divisions of the autonomic nervous system, the others being the sympathetic nervous system and the enteric nervous system. The enteric nervous system is sometimes considered part of t ...
division of the
autonomic nervous system The autonomic nervous system (ANS), formerly referred to as the vegetative nervous system, is a division of the peripheral nervous system that supplies viscera, internal organs, smooth muscle and glands. The autonomic nervous system is a control ...
with minimal input from the central nervous system. Parasympathetic branches extend from the sacral plexus into the arteries supplying the erectile tissue; upon stimulation, these nerve branches release
acetylcholine Acetylcholine (ACh) is an organic chemical that functions in the brain and body of many types of animals (including humans) as a neurotransmitter. Its name is derived from its chemical structure: it is an ester of acetic acid and choline. Part ...
, which in turn causes release of
nitric oxide Nitric oxide (nitrogen oxide or nitrogen monoxide) is a colorless gas with the formula . It is one of the principal oxides of nitrogen. Nitric oxide is a free radical: it has an unpaired electron, which is sometimes denoted by a dot in its che ...
from
endothelial cell The endothelium is a single layer of squamous endothelial cells that line the interior surface of blood vessels and lymphatic vessels. The endothelium forms an interface between circulating blood or lymph in the lumen and the rest of the vessel ...
s in the
trabecular A trabecula (plural trabeculae, from Latin for "small beam") is a small, often microscopic, tissue element in the form of a small beam, strut or rod that supports or anchors a framework of parts within a body or organ. A trabecula generally has ...
arteries. Nitric oxide diffuses to the
smooth muscle Smooth muscle is an involuntary non-striated muscle, so-called because it has no sarcomeres and therefore no striations (''bands'' or ''stripes''). It is divided into two subgroups, single-unit and multiunit smooth muscle. Within single-unit mus ...
of the arteries (called ''trabecular smooth muscle''), acting as a vasodilating agent. The arteries dilate, filling the ''corpus spongiosum'' and ''corpora cavernosa'' with blood. The ischiocavernosus and
bulbospongiosus The bulbospongiosus muscle (bulbocavernosus in older texts) is one of the superficial muscles of the perineum. It has a slightly different origin, insertion and function in males and females. In males, it covers the bulb of the penis. In femal ...
muscles also compress the veins of the corpora cavernosa, limiting the venous drainage of blood. Erection subsides when parasympathetic stimulation is discontinued; baseline stimulation from the
sympathetic division The sympathetic nervous system (SNS) is one of the three divisions of the autonomic nervous system, the others being the parasympathetic nervous system and the enteric nervous system. The enteric nervous system is sometimes considered part of the ...
of the autonomic nervous system causes constriction of the penile arteries and cavernosal sinosoids, forcing blood out of the erectile tissue via erection-related veins which include one deep dorsal vein, a pair of cavernosal veins, and two pairs of para-arterial veins between Buck's fascia and the tunica albuginea. Erection rigidity is mechanically controlled by reduction blood flow via theses veins, and thereby building up the pressure of the corpus cavernosum and corpus spongiosum, an integral instructure, the distal ligament, buttresses the glans penis. After ejaculation or cessation of stimulation, erection usually subsides, but the time taken may vary depending on the length and thickness of the penis.


Voluntary and involuntary control

The cerebral cortex can initiate erection in the absence of direct mechanical stimulation (in response to visual, auditory, olfactory, imagined, or tactile stimuli) acting through erectile centers in the lumbar and sacral regions of the spinal cord. The cortex may suppress erection, even in the presence of mechanical stimulation, as may other psychological, emotional, and environmental factors.


Nocturnal erection

The penis may become erect during sleep or be erect on waking up. Such an erection is medically known as nocturnal penile tumescence (informally: ''morning wood'' or ''morning glory'').


Urination

Urinating with an erection can be more difficult than without. David Samadi, director of men's health at St. Francis Hospital in
Long Island Long Island is a densely populated island in the southeastern region of the U.S. state of New York (state), New York, part of the New York metropolitan area. With over 8 million people, Long Island is the most populous island in the United Sta ...
, says "To prevent semen from entering into the bladder, the internal urethral sphincter contracts." This also works in opposite way, making it less likely that urine will pass through the urethra with an erect penis. Combined with morning wood, and a full bladder from overnight, many people may have experience with the extra challenge that comes with the combination. Medical recommendations include: # Waiting for the erection to go away, including techniques for relaxing # Peeing while bending over or sitting down # Massaging, or putting light pressure or a hot compress on the bladder (from the outside, between the navel and the pubic bone)


Socio-sexual aspects


Social

Though an erection can have many causes, the more common indicator of
sexual arousal Sexual arousal (also known as sexual excitement) describes the physiological and psychological responses in preparation for sexual intercourse or when exposed to sexual stimuli. A number of physiological responses occur in the body and mind as ...
has a prevalent view over society and its portrayal in public is considered taboo in many societies, probably lesser than public sex but higher than
nudity Nudity is the state of being in which a human is without clothing. The loss of body hair was one of the physical characteristics that marked the biological evolution of modern humans from their hominin ancestors. Adaptations related to ...
. Erectile dysfunction is considered a flaw that associates shame at the individual and his partner. The penile plethysmograph, which measures erections, has been used by some governments and courts of law to measure sexual orientation. An unusual aversion to the erect penis is sometimes referred to as phallophobia.


Spontaneous or random erections

Spontaneous erections, also known as involuntary, random or unwanted erections, are commonplace and a normal part of male physiology. Socially, such erections can be
embarrassing Embarrassment or awkwardness is an emotional state that is associated with mild to severe levels of discomfort, and which is usually experienced when someone commits (or thinks of) a socially unacceptable or frowned-upon act that is witnessed ...
if they happen in public or when undesired. Such erections can occur at any time of day, and if clothed may cause a bulge which (if required) can be disguised or hidden by wearing close-fitting underwear, a long shirt, or baggier clothes. Few slang terms associated with erect penis outline becoming visible from underneath clothing are ''mooseknuckle'' and ''manbulge''.


Size

The length of the flaccid penis is not necessarily indicative of the length of the penis when it becomes erect, with some smaller flaccid penises growing much longer, and some larger flaccid penises growing comparatively less. Generally, the size of an erect penis is fixed throughout post-
pubescent The adjective pubescent may describe: * people or animals undergoing puberty * plants that are hairy, covered in trichomes * insects that are covered in setae In biology, setae (singular seta ; from the Latin word for "bristle") are any of a ...
life. Its size may be increased by surgery, although penile enlargement is controversial, and a majority of men were "not satisfied" with the results, according to one study. Though the size of a penis varies considerably between males, the average length of an erect human penis is 13.12 cm (5.17 inches), while the average circumference of an erect human penis is 11.66 cm (4.59 inches).


Direction

Although many erect penises point upwards, it is common and normal for the erect penis to point nearly vertically upwards or horizontally straight forward or even nearly vertically downwards, all depending on the tension of the suspensory ligament that holds it in position. An erect penis can also take on a number of different shapes, ranging from a straight tube to a tube with a curvature up or down or to the left or right. An increase in penile curvature can be caused by Peyronie's disease. This may cause physical and psychological effects for the affected individual, which could include
erectile dysfunction Erectile dysfunction (ED), also called impotence, is the type of sexual dysfunction in which the penis fails to become or stay erect during sexual activity. It is the most common sexual problem in men.Cunningham GR, Rosen RC. Overview of male ...
or pain during an erection. Treatments include oral medication (such as colchicine) or surgery, which is most often performed only as a last resort. The following table shows how common various erection angles are for a standing male. In the table, zero degrees (0°) is pointing straight up against the abdomen, 90° is horizontal and pointing straight forward, and 180° is pointing straight down to the feet. An upward pointing angle is most common and the average erection angle is 74.3 degrees. The penile curvature was measured same time. 63% men have straight penis. 22.2% men have upwards curvature and 14.8% men have downwards curvature.


Medical conditions


Erectile dysfunction

Erectile dysfunction (also known as ED or "(male) impotence") is a sexual dysfunction characterized by the inability to develop and/or maintain an erection. The study of erectile dysfunction within medicine is known as andrology, a sub-field within urology. Erectile dysfunction may occur due to
physiological Physiology (; ) is the scientific study of functions and mechanisms in a living system. As a sub-discipline of biology, physiology focuses on how organisms, organ systems, individual organs, cells, and biomolecules carry out the chemical ...
or
psychological Psychology is the scientific study of mind and behavior. Psychology includes the study of conscious and unconscious phenomena, including feelings and thoughts. It is an academic discipline of immense scope, crossing the boundaries between t ...
reasons, most of which are amenable to treatment. Common physiological reasons include diabetes, kidney disease, chronic alcoholism, multiple sclerosis, atherosclerosis, vascular disease, including arterial insufficiency and venogenic erectile dysfunction, and neurologic disease which collectively account for about 70% of ED cases. Some drugs used to treat other conditions, such as lithium and paroxetine, may cause erectile dysfunction. Erectile dysfunction, tied closely as it is to cultural notions of
potency Potency may refer to: * Potency (pharmacology), a measure of the activity of a drug in a biological system * Virility * Cell potency, a measure of the differentiation potential of stem cells * In homeopathic dilutions, potency is a measure of how ...
,
success Success is the state or condition of meeting a defined range of expectations. It may be viewed as the opposite of failure. The criteria for success depend on context, and may be relative to a particular observer or belief system. One person mig ...
and
masculinity Masculinity (also called manhood or manliness) is a set of attributes, behaviors, and roles associated with men and boys. Masculinity can be theoretically understood as socially constructed, and there is also evidence that some behaviors con ...
, can have devastating psychological consequences including feelings of shame, loss or inadequacy. There is a strong
culture of silence A conspiracy of silence, or culture of silence, describes the behavior of a group of people of some size, as large as an entire national group or profession or as small as a group of colleagues, that by unspoken consensus does not mention, discuss ...
and inability to discuss the matter. Around one in ten men experience recurring impotence problems at some point in their lives.


Priapism

Priapism Priapism is a condition in which a penis remains erect for hours in the absence of stimulation or after stimulation has ended. There are three types: ischemic (low-flow), nonischemic (high-flow), and recurrent ischemic (intermittent). Most cases ...
is a painful condition in which the penis does not return to its flaccid state, despite the absence of both physical and psychological stimulation. Priapism lasting over four hours is a medical emergency.


In non-human animals

At the time of penetration, the canine penis is not erect, and only able to penetrate the female because it includes a narrow bone called the
baculum The baculum (also penis bone, penile bone, or ''os penis'', ''os genitale'' or ''os priapi'') is a bone found in the penis of many placental mammals. It is absent from the human penis, but present in the penises of some primates, such as the ...
, a feature of most placental mammals. After the male achieves penetration, he will often hold the female tighter and thrust faster, and it is during this time that the male's penis expands. Unlike human sexual intercourse, where the male penis commonly becomes erect before entering the female, canine copulation involves the male first penetrating the female, after which swelling of the penis to erection occurs. An elephant's penis is S-shaped when fully erect and has a Y-shaped orifice. Given the small amount of erectile tissue in a
bull's penis A bull is an intact (i.e., not castrated) adult male of the species ''Bos taurus'' (cattle). More muscular and aggressive than the females of the same species (i.e., cows), bulls have long been an important symbol in many religions, includin ...
, there is little enlargement after erection. The penis is quite rigid when non-erect, and becomes even more rigid during erection. Protrusion is not affected much by erection, but more by relaxation of the retractor penis muscle and straightening of the sigmoid flexure. A male fossa's penis reaches to between his forelegs when erect. When not erect, a
horse's penis A stallion is a male horse that has not been gelded (castrated). Stallions follow the conformation and phenotype of their breed, but within that standard, the presence of hormones such as testosterone may give stallions a thicker, "cresty" neck ...
is housed within the prepuce, long and in diameter with the distal end . The
retractor muscle Retractor may refer to: * Retractor (medical), a medical instrument * Retractor (memory), a person * Retractor (chess), type of a chess problem * Retractor muscle Skeletal muscles (commonly referred to as muscles) are organs of the vertebrate ...
contracts to retract the penis into the sheath and relaxes to allow the penis to extend from the sheath. When erect, the penis doubles in length and thickness and the glans increases by 3 to . Erection and protrusion take place gradually, by the increasing tumescence of the erectile vascular tissue in the '' corpus cavernosum penis''. Most stallions achieve erection within 2 minutes of contact with an estrus mare, and mount the estrus mare 5–10 seconds afterward. A bird penis is different in structure from mammal penises, being an erectile expansion of the cloacal wall and being erected by lymph, not blood. The penis of the
lake duck The lake duck (''Oxyura vittata'') is a small, South American stiff-tailed duck. It is also called the Argentine blue-bill, Argentine blue-billed duck, Argentine lake duck, or Argentine ruddy duck. Description The lake duck grows to . Females ...
can reach about the same length as the animal himself when fully erect, but more commonly is about half the bird's length.


Terminology

Clinically, erection is often known as "penile erection", and the state of being erect, and process of erection, are described as "tumescence" or "penile tumescence". The term for the subsiding or cessation of an erection is "
detumescence Tumescence is the quality or state of being tumescent or swollen. Tumescence usually refers to the normal engorgement with blood (vascular congestion) of the erectile tissues, marking sexual excitation, and possible readiness for sexual activity. T ...
". Colloquially and in slang, erection is known by many informal terms. Commonly encountered English terms include 'stiffy', 'hard-on', 'boner' and 'woody'. There are several slang words, euphemisms and synonyms for an erection in English and in other languages. (See also The WikiSaurus entry.)


See also

* Clitoral erection * Cock ring *
Death erection A death erection, angel lust, rigor erectus, or terminal erection is a post-mortem erection, technically a priapism, observed in the corpses of men who have been executed, particularly by hanging. "Erection has long been observed to follow injuri ...
* Human penis * Issues in social nudity * Nipple stimulation * Nocturnal penile tumescence *
Priapism Priapism is a condition in which a penis remains erect for hours in the absence of stimulation or after stimulation has ended. There are three types: ischemic (low-flow), nonischemic (high-flow), and recurrent ischemic (intermittent). Most cases ...
*
Sexual function Sexual function is how the body reacts in different stages of the sexual response cycle, or as a result of sexual dysfunction. Relevant aspects of sexual function are defined on the basis of a modified version of Masters and Johnson's work. The aspe ...


References

{{Authority control Sexual arousal Andrology Men's health