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The Epipaschiinae are a
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
of
snout moth The Pyralidae, commonly called pyralid moths, snout moths or grass moths, are a family of Lepidoptera in the ditrysian superfamily Pyraloidea. In many (particularly older) classifications, the grass moths (Crambidae) are included in the Pyralida ...
s (
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Pyralidae). More than 720 species are known today, which are found mainly in the tropics and subtropics. Some occur in temperate regions, but the subfamily is apparently completely absent from Europe, at least as native species. A few Epipaschiinae are crop pests that may occasionally become economically significant.


Description and ecology

Adult females are often hard to distinguish from related lineages, and even the larvae do not possess the characteristic sclerotized bristle base near the start of the abdomen, whose position is a tell-tale mark of the other subfamilies of Pyralidae. By contrast, the adult males of Epipaschiinae are easier to recognize, and three of their traits support the assumption that Epipaschiinae are a natural, monophyletic group: # an always upturned and pointed third segment of the labial palpi # a ventrally curved phallobase of the male which usually extends beyond the ductus ejaculatorius # the weakly sclerotized tegumen In addition, in most cases the adult males of this subfamily have a conspicuous scaled projection from the scape of the antennae. The
caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder Sym ...
larvae are leaf rollers, leaf tiers and
leaf miner A leaf miner is any one of numerous species of insects in which the larval stage lives in, and eats, the leaf tissue of plants. The vast majority of leaf-mining insects are moths (Lepidoptera), sawflies (Symphyta, the mother clade of wasps), ...
s. As pests, they infest such diverse plants as ''
Persea americana The avocado (''Persea americana'') is a medium-sized, evergreen tree in the laurel family ( Lauraceae). It is native to the Americas and was first domesticated by Mesoamerican tribes more than 5,000 years ago. Then as now it was prized for ...
'' (avocado), ''
Swietenia ''Swietenia'' is a genus of trees in the chinaberry family, Meliaceae. It occurs natively in the Neotropics, from southern Florida, the Caribbean, Mexico and Central America south to Bolivia. The genus is named for Dutch-Austrian physician Ge ...
'' (mahoganies), or ''
Zea mays Maize ( ; ''Zea mays'' subsp. ''mays'', from es, maíz after tnq, mahiz), also known as corn (North American and Australian English), is a cereal grain first domesticated by indigenous peoples in southern Mexico about 10,000 years ago. Th ...
'' (corn). However, they are usually a mere nuisance and do not cause large-scale crop failure.


Systematics

Alma Solis (1993) provided a phylogenetic analysis of 20 genera of the ''Pococera'' complex, consisting of some 300 species in the Western Hemisphere. *'' Accinctapubes'' Solis, 1993 *'' Agastophanes'' Turner, 1937 *'' Anaeglis'' Lederer, 1863 *'' Anarnatula'' Dyar, 1918 *'' Anexophana'' Viette, 1960 *'' Apocera'' Schaus, 1912 (= ''Paranatula'' Dyar, 1913) *'' Araeopaschia'' Hampson, 1906 (= ''Aroeopaschia'' Amsel, 1956) *'' Astrapometis'' Meyrick, 1884 *'' Austropaschia'' Hampson, 1916 *'' Axiocrita'' Turner, 1913 *'' Bibasilaris'' Solis, 1993 *''
Cacozelia ''Cacozelia'' is a genus of snout moths. It was described by Augustus Radcliffe Grote in 1878 and is known from Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), ...
'' Grote, 1878 *'' Calybitia'' Schaus, 1922 *'' Canipsa'' Walker, 1866 (= ''Sarama'' Moore, 1888, ''Scopocera'' Moore, 1888) *'' Carthara'' Walker, 1865 (= ''Leptosphetta'' Butler, 1878, ''Pycnulia'' Zeller, 1881) *'' Catalaodes'' Viette, 1953 *'' Catamola'' Meyrick, 1884 (= ''Elaphernis'' Meyrick, 1936) *'' Cecidipta'' Berg, 1877 (= ''Acecidipta'' Amsel, 1956) *'' Chloropaschia'' Hampson, 1906 *'' Coenodomus'' Walsingham, 1888 (= ''Alippa'' Aurivillius, 1894, ''Dyaria'' Neumoegen, 1893) *'' Dasyvesica'' Solis, 1991 *'' Deuterollyta'' Lederer, 1863 (= ''Ajacania'' Schaus, 1925, ''Ajocara'' Schaus, 1925, ''Oedomia'' Dognin, 1906, ''Winona'' Hulst, 1888) *'' Doddiana'' Turner, 1902 *'' Elisabethinia'' Ghesquière, 1942 *'' Ephedrophila'' Dumont, 1928 *'' Epilepia'' Janse, 1931 *'' Epipaschia'' Clemens, 1860 *'' Eublemmodes'' Gaede, 1917 *'' Geropaschia'' Hampson, 1917 (= ''Araeopaschia'' Hampson, 1916) *'' Glossopaschia'' Dyar, 1914 *'' Heminomistis'' Meyrick, 1933 *'' Homura'' Lederer, 1863 *'' Incarcha'' Dyar, 1910 *'' Isolopha'' Hampson, 1895 (= ''Islopha'' Janse, 1931) *''
Jocara ''Jocara'' is a genus of snout moths. It was described by Francis Walker in 1863. Species * '' Jocara abachuma'' * '' Jocara aidana'' * '' Jocara albiferalis'' * '' Jocara albimedialis'' * '' Jocara albimedialis'' * '' Jocara amazonalis'' * '' ...
'' Walker, 1863 (= '' Toripalpus'' Grote, 1878) *'' Lacalma'' Janse, 1931 *'' Lameerea'' Ghesquière, 1942 *'' Lamida'' Walker, 1859 (= ''Allata'' Walker, 1863) *'' Lepidogma'' Meyrick, 1890 (= ''Asopina'' Christoph, 1893, ''Precopia'' Ragonot, 1891) *'' Lepipaschia'' J. C. Shaffer & Solis, 1994 *'' Leptoses'' Ghesquière, 1942 *''
Lista Lista is a former municipality located in the old Vest-Agder county in Norway. The municipality existed from 1838 until its dissolution in 1965. The administrative centre was the village of Vanse where Vanse Church is located. Lista municipa ...
'' Walker, 1859 (= ''Belonepholis'' Butler, 1889, ''Craneophora'' Christoph, 1881, ''Paracme'' Lederer, 1863) *'' Locastra'' Walker, 1859 (= ''Taurica'' Walker, 1866) *''
Macalla ''Macalla'' is a 1985 musical album by Irish folk group Clannad. It is their ninth album and became prominent with a collaboration between Clannad's singer Moya Brennan and U2 vocalist Bono on the duet "In a Lifetime". Furthermore, it feature ...
'' Walker, 1859 (= ''Aradrapha'' Walker, 1866, ''Mochlocera'' Grote, 1876, ''Pseudomacalla'' Dognin, 1908) *'' Mazdacis'' Solis, 1993 *'' Mediavia'' Solis, 1993 *'' Micropaschia'' Hampson, 1906 *'' Milgithea'' Schaus, 1922 (= ''Miligithea'' Neave, 1940) *'' Mimaglossa'' Warren, 1891 *'' Neopaschia'' Janse, 1922 *'' Noctuides'' Staudinger, 1892 (= ''Anartula'' Staudinger, 1893, ''Arnatula'' Hampson, 1896, ''Parorthaga'' Hampson, 1896) *'' Nouanda'' Holland & Schaus, 1925 *'' Nyctereutica'' Turner, 1904 (= ''Diastrophica'' Turner, 1937) *'' Obutobea'' Ghesquière, 1942 *'' Odontopaschia'' Hampson, 1903 *'' Omphalepia'' Hampson, 1906 *'' Omphalota'' Hampson, 1899 *''
Oneida Oneida may refer to: Native American/First Nations * Oneida people, a Native American/First Nations people and one of the five founding nations of the Iroquois Confederacy * Oneida language * Oneida Indian Nation, based in New York * Oneida Na ...
'' Hulst, 1889 *'' Orthaga'' Walker, 1859 (= ''Edeta'' Walker, 1859, ''Hyperbalanotis'' Warren, 1891, ''Pannucha'' Moore, 1888, ''Proboscidophora'' Warren, 1891) *'' Oxyalcia'' Dognin, 1905 *'' Pandoflabella'' Solis, 1993 *'' Parastericta'' Janse, 1931 *'' Peplochora'' Meyrick, 1933 *'' Phidotricha'' Ragonot, 1889 (= ''Eutrichocera'' Hampson, 1904, ''Jocarula'' Dyar, 1925) *'' Plumiphora'' Janse, 1931 *'' Plutopaschia'' Hampson, 1917 *'' Pococera'' Zeller, 1848 (= ''Attacapa'' Hulst, 1889, ''Auradisa'' Walker, 1866, ''Benta'' Walker, 1863, ''Hemimatia'' Lederer, 1863, ''Katona'' Hulst, 1888, ''Lanthaphe'' Clemens, 1860, ''Lanthape'' Hulst, 1903, ''Loma'' Hulst, 1888, ''Saluda'' Hulst, 1888, ''Tetralopha'' Zeller, 1848, ''Tioga'' Hulst, 1888, ''Afra'' Ghesquière, 1942, ''Wanda'' Hulst, 1888) *'' Poliopaschia'' Hampson, 1916 *'' Polylophota'' Hampson, 1906 *'' Pseudocera'' Walker, 1863 *'' Quadraforma'' Solis, 1993 *'' Rhynchopaschia'' Hampson, 1906 *'' Roeseliodes'' Warren, 1891 *''
Salma The South American land mammal ages (SALMA) establish a geologic timescale for prehistoric South American fauna beginning 64.5 Ma during the Paleocene and continuing through to the Late Pleistocene (0.011 Ma). These periods are referred to as a ...
'' Walker, 1863 (= ''Calinipaxa'' Walker, 1866, ''Enchesphora'' Turner, 1913, ''Exacosmia'' Walker, 1865, ''Heterobella'' Turner, 1904, ''Heterobela'' Turner, 1904, ''Orthotrichophora'' Warren, 1891, ''Parasarama'' Warren, 1890, ''Parasamera'' Sharp, 1892, ''Pseudolocastra'' Snellen, 1890, ''Pseudolocastra'' Warren, 1891) *'' Schoutedenidea'' Ghesquière, 1942 *'' Sparactica'' Meyrick, 1938 *'' Spectrotrota'' Warren, 1891 *'' Speiroceras'' Chrétien, 1911 *'' Stericta'' Lederer, 1863 (= ''Glossina'' Guenée, 1854, ''Matalia'' Walker, 1866, ''Oncobela'' Turner, 1937, ''Phialia'' Walker, 1866) *'' Sultania'' Koçak, 1987 *'' Taiwanastrapometis'' Shibuya, 1928 *'' Tallula'' Hulst, 1888 *'' Tancoa'' Schaus, 1922 *'' Teliphasa'' Moore, 1888 (= ''Sultania'' Koçak, 1987) *'' Termioptycha'' Meyrick, 1889 (= ''Sialocyttara'' Turner, 1913) *'' Tineopaschia'' Hampson, 1916 *'' Titanoceros'' Meyrick, 1884 *'' Trichotophysa'' Warren, 1896 *'' Yuma'' Hulst, 1889


References

* (2011)
Epipaschiinae Meyrick
''Lepidoptera and Some Other Life Forms''. Retrieved May 12, 2017. * (2007): Phylogenetic studies and modern classification of the Pyraloidea (Lepidoptera). '' Revista Colombiana de Entomología'' 33(1): 1–8 nglish with Spanish abstractbr>HTML fulltext


External links



at the University of Florida Featured Creatures website

at the University of Florida Featured Creatures website {{Taxonbar, from=Q3055773 Moth subfamilies