Epinephelus Rivulatus Juvénile à Confirmer
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''Epinephelus'' is a genus of marine ray-finned fish, groupers from the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
Epinephelinae, part of the family Serranidae, which also includes the
anthias Anthias are members of the family Serranidae and make up the subfamily Anthiinae. Anthias make up a sizeable portion of the population of pink, orange, and yellow reef fishes seen swarming in most coral reef photography and film. The name Anthi ...
and sea basses. They are predatory fish, largely associated with reefs and are found in tropical and subtropical seas throughout the world. They are important target species for fisheries.


Characteristics

The fishes in the genus ''Epinephelus'' have elongate, subcylindrical bodies which may be oblong or deep and compressed in shape. The depth of the body varies between 2.3 and 3.7 times the
standard length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish m ...
and head is usually around the same length as the body is deep. The preopercle can be rounded or angular and has a serrated rear edge with the serrations at the angle being enlarged to a lesser or greater extent. In a small number of species serrations are small and on the lower edge they are covered by skin. Caniform teeth are found at the front of jaws, although these can be rather small in a few species. They do not have any obviously enlarged caniform teeth in the middle of the lower jaw. There are teeth on the roof of the mouth. In adults, the maxilla does not have a noticeable bony protrusion on the lower rear angle, although they can have an deep step or hook-like process which is hidden by the upper lips, on the rear part of its lower edge. The dorsal fin normally contains 9 spines, although some species have 10, as well as 12 to 19 rays. The origin of the dorsal fin sits above the opercle and the soft rayed part is shorter than the spiny part. The
anal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
contains 3 distinct spines and 7 to 10 soft rays. The pectoral fin is rounded with its middle rays being longer than the others longest. The caudal fin may be rounded, truncate or concave, contains 8 branched rays and 8 to 10 fin rays which are slender, unbranched and unsegmented (referred to as "procurrent") fin rays at the leading edges of he caudal fin on the upper lobe and 7 branched rays and 7 to 10 procurrent rays in the lower lobe. The body is covered in ctenoid or smooth scales.


Habitat and biology

''Epinephelus'' groupers are occur mainly on coral or rocky reefs, although a small number of species have been recorded over substrates consisting of sand, silt or mud. A few species are found in deep water, down to at least , but the majority occur between . The two largest members of the genus, '' E. itajara'' and '' E. lanceolatus'', either of which may attain a length in excess of and a weight greater than have frequently been recorded in estuaries and harbours. Most of the species in the genus ''Epinephelus'' are predatory fish which feed on larger invertebrates, mostly crustaceans, and other fishes taken on or close to the substrate. E. undulosus is an unusual grouper species distinguished by having many, long gill rakers and this species has been reported to feed on pelagic tunicates, at least on occasion. Only a few species have had their reproductive biology studied and many species appear to be protogynous hermaphrodites. However, in some species there are males in the populations which are smaller than some of the females, suggesting a more complex biology and this suggests that some females do not change sex, and that some males may not have a undergone a functional female stage.


Distribution

''Epinephelus'' species are found around the world in tropical and subtropical seas and oceans. The greatest diversity occurs in the Indo-West Pacific, while 8 species are found in the eastern Pacific, 11 in the western Atlantic Ocean 9 species in the eastern Atlantic Ocean and the Mediterranean. Four species have entered the Mediterranean Sea from the Red Sea via the
Suez Canal The Suez Canal ( arz, قَنَاةُ ٱلسُّوَيْسِ, ') is an artificial sea-level waterway in Egypt, connecting the Mediterranean Sea to the Red Sea through the Isthmus of Suez and dividing Africa and Asia. The long canal is a popular ...
as Lessepsian migrants.


Utilisation

''Epinephelus'' groupers are among the most valuable species exploited by commercial fishes in the world's tropical seas and they fetch some of the highest prices when marketed. They have also been used in
aquaculture Aquaculture (less commonly spelled aquiculture), also known as aquafarming, is the controlled cultivation ("farming") of aquatic organisms such as fish, crustaceans, mollusks, algae and other organisms of value such as aquatic plants (e.g. lot ...
.


Species

The 89 recognized species in this genus are: Some of these species are placed in the genus '' Hyporthodus'' by some authorities, for example ''Epinephelus darwinensis'' is treated as ''Hyporthodus darwinensis'' by the '' Catalog of Fishes''.


References


External links

* * {{Taxonbar, from=Q284420 Epinephelini Extant Eocene first appearances Taxa named by Marcus Elieser Bloch