Epinephelus Morio
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The red grouper (''Epinephelus morio'') is a species of marine
ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or h ...
, a
grouper Groupers are fish of any of a number of genera in the subfamily Epinephelinae of the family Serranidae, in the order Perciformes. Not all serranids are called "groupers"; the family also includes the sea basses. The common name "grouper" is ...
from the subfamily
Epinephelinae Groupers are fish of any of a number of genera in the subfamily Epinephelinae of the family Serranidae, in the order Perciformes. Not all serranids are called "groupers"; the family also includes the sea basses. The common name "grouper" is ...
which is part of the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Serranidae The Serranidae are a large family of fishes belonging to the order Perciformes. The family contains about 450 species in 65 genera, including the sea basses and the groupers (subfamily Epinephelinae). Although many species are small, in some ca ...
, which also includes the
anthias Anthias are members of the family Serranidae and make up the subfamily Anthiinae. Anthias make up a sizeable portion of the population of pink, orange, and yellow reef fishes seen swarming in most coral reef photography and film. The name Anthi ...
and sea basses. It is found in the western Atlantic Ocean.


Description

The red grouper has a body with a standard length which is 2.6 to 3 times as long as it is deep. The preopercle is subangular with the serrations at its angle being slightly enlarged and the upper edge of the gill cover is straight. The gill cover has three flat spines with the central spine being the longest. The
dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through c ...
contains 11 spines and 16-17 soft rays while the anal fin has 3 spines and 8 soft rays. The
pectoral fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as ...
s are longer than the
pelvic fins Pelvic fins or ventral fins are paired fins located on the ventral surface of fish. The paired pelvic fins are homologous to the hindlimbs of tetrapods. Structure and function Structure In actinopterygians, the pelvic fin consists of two e ...
and the
caudal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as ...
is truncate. The They are dark reddish brown on the upper part of the head and body, shading to paler pink on the underparts, they are marked with lighter spots and blotches across their body and there are darker margins to the fins. This species has a maximum published
total length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish ...
of , although they a more commonly found at lengths around , and a maximum published weight of .


Distribution

The red grouper's typical range is coastal areas in the western Atlantic, stretching from southern Brazil to North Carolina in the US and including the Gulf of Mexico and
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.


Habitat and biology

The red grouper is a demersal, largely sedentary species which has an extended (~40 day) pelagic larval stage before it settles in shallow coastal hardbottom habitat as juveniles. They remain in inshore waters for 4–5 years before migrating to offshore hardbottom habitat—particularly on the edge of the continental self—as adults. Spawning occurs offshore between January and June, peaking in May. While primarily eating benthic invertebrates, the red grouper is an opportunistic feeder in the reef community. The diet commonly includes xanthid and portunid crabs, juvenile spiny lobster, and snapping shrimp, with the occasional fish. The red grouper is of moderate size, about 125 cm and weighs 23 kg or more. Body coloration is typically reddish-brown in color often, with many white spots. When aggravated (they are highly territorial) or involved in spawning activities, these fish can very rapidly change coloration patterns, with the head or other parts of the body turning completely white, and the white spots appearing more intense.


Habitat engineers of the sea

Red grouper actively excavate pits in the seafloor. They start digging in the sediment from the time they settle out of the plankton and continue throughout their lifetime. They use their caudal fin and their mouths to remove debris and sediment from rocks, creating exposed surfaces on which sessile organisms actively settle (e.g., sponges, soft corals, algae). The exposure of structure also attracts a myriad of other species, including mobile invertebrates and a remarkable diversity of other fishes, from gobies and butterflyfish to grunts and snapper. The
lionfish ''Pterois'' is a genus of venomous marine fish, commonly known as lionfish, native to the Indo-Pacific. Also called firefish, turkeyfish, tastyfish, or butterfly-cod, it is characterized by conspicuous warning coloration with red, white, crea ...
''Pterois volitans'' started invading red grouper habitat by 2008, from Florida Bay to the Florida Keys and offshore to
Pulley Ridge Pulley Ridge is a mesophotic coral reef system off the shores of the continental United States. The reef rests on sunken barrier islands and lies 100 miles west of the Tortugas Ecological Reserve and stretches north about 60 miles at depths rang ...
, a mesophotic coral reef on the West Florida Shelf west of the Dry Tortugas. Known for being extremely capable predators on small reef fish, scientists are very interested in determining the extent to which their invasion changes the functional dynamics of associated communities.


Parasites

As other fish, red groupers harbour a number of
parasites Parasitism is a close relationship between species, where one organism, the parasite, lives on or inside another organism, the host, causing it some harm, and is adapted structurally to this way of life. The entomologist E. O. Wilson ha ...
, including, on its
gills A gill () is a respiratory organ that many aquatic organisms use to extract dissolved oxygen from water and to excrete carbon dioxide. The gills of some species, such as hermit crabs, have adapted to allow respiration on land provided they are ...
, the
monogenea Monogeneans are a group of ectoparasitic flatworms commonly found on the skin, gills, or fins of fish. They have a direct lifecycle and do not require an intermediate host. Adults are hermaphrodites, meaning they have both male and female repro ...
ns '' Pseudorhabdosynochus justinella'' and '' Pseudorhabdosynochus yucatanensis''.


Taxonomy

The red grouper was first formally described as ''Serranus morio'' in 1828 by the French
zoologist Zoology ()The pronunciation of zoology as is usually regarded as nonstandard, though it is not uncommon. is the branch of biology that studies the animal kingdom, including the structure, embryology, evolution, classification, habits, and d ...
Achille Valenciennes (1794-1865), with the type locality given as the fish market in New York, the fish being "probably caught south of there".


Utilisation

The red grouper is a commercially important species for fisheries throughout its range and it is also an valuable resource for recreational fisheries too.


References


External links

* {{Taxonbar, from=Q2414243 Epinephelus Fish of Guatemala Fish of the Atlantic Ocean Sport fish Fish described in 1828 Taxonomy articles created by Polbot