An epicotyl is important for the beginning stages of a plant's life. It is the region of a seedling stem above the stalks of the seed leaves of an embryo plant. It grows rapidly, showing hypogeal germination, and extends the stem above the soil surface. A common misconception is that the epicotyl, being closer to the apex of the plant, is the first part to emerge after germination - rather, the hypocotyl, the region of the stem between the point of attachment of the cotyledons and the root - forms a hook during hypogeal germination and pushes out of the soil, allowing the more delicate tissues of the plumules and apical meristem to avoid damage from pushing through the soil. The epicotyl will expand and form the point of attachment of the shoot apex and leaf primordia or "first true leaves". Cotyledons may remain belowground or be pushed up aboveground with the growing stem depending on the plant species in question.
In
plant physiology
Plant physiology is a subdiscipline of botany concerned with the functioning, or physiology, of plants. Closely related fields include plant morphology (structure of plants), plant ecology (interactions with the environment), phytochemistry (bi ...
, the epicotyl is the embryonic shoot above the
cotyledon
A cotyledon (; ; ; , gen. (), ) is a significant part of the embryo within the seed of a plant, and is defined as "the embryonic leaf in seed-bearing plants, one or more of which are the first to appear from a germinating seed." The numb ...
s. In most plants the epicotyl will eventually develop into the leaves of the plant. In
dicots, the
hypocotyl
The hypocotyl (short for "hypocotyledonous stem", meaning "below seed leaf") is the stem of a germinating seedling, found below the cotyledons (seed leaves) and above the radicle (root).
Eudicots
As the plant embryo grows at germination, it send ...
is what appears to be the base stem under the spent withered cotyledons, and the shoot just above that is the epicotyl. In
monocot
Monocotyledons (), commonly referred to as monocots, (Lilianae ''sensu'' Chase & Reveal) are grass and grass-like flowering plants (angiosperms), the seeds of which typically contain only one embryonic leaf, or cotyledon. They constitute one of ...
plants, the first shoot that emerges from the ground or from the
seed
A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...
is the epicotyl, from which the first shoots and leaves emerge.
Lengthening of the epicotyl is thought to be controlled by the phytochrome photoreceptors.
See also
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Hypocotyl
The hypocotyl (short for "hypocotyledonous stem", meaning "below seed leaf") is the stem of a germinating seedling, found below the cotyledons (seed leaves) and above the radicle (root).
Eudicots
As the plant embryo grows at germination, it send ...
*
Radicle
In botany, the radicle is the first part of a seedling (a growing plant embryo) to emerge from the seed during the process of germination. The radicle is the embryonic root of the plant, and grows downward in the soil (the shoot emerges from ...
*
Plumule
A seedling is a young sporophyte developing out of a plant embryo from a seed. Seedling development starts with germination of the seed. A typical young seedling consists of three main parts: the radicle (embryonic root), the hypocotyl (embr ...
External links
* "epicotyl." A Dictionary of Biology. 2004. Encyclopedia.com. 3 Dec. 2015
Plant morphology