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Entomology () is the scientific study of insects, a branch of zoology. In the past the term "insect" was less specific, and historically the definition of entomology would also include the study of animals in other arthropod groups, such as arachnids, myriapods, and crustaceans. This wider meaning may still be encountered in informal use. Like several of the other fields that are categorized within zoology, entomology is a taxon-based category; any form of scientific study in which there is a focus on insect-related inquiries is, by definition, entomology. Entomology therefore overlaps with a cross-section of topics as diverse as molecular genetics,
behavior Behavior (American English) or behaviour (British English) is the range of actions and mannerisms made by individuals, organisms, systems or artificial entities in some environment. These systems can include other systems or organisms as wel ...
, neuroscience, biomechanics, biochemistry,
systematics Biological systematics is the study of the diversification of living forms, both past and present, and the relationships among living things through time. Relationships are visualized as evolutionary trees (synonyms: cladograms, phylogenetic tre ...
, physiology,
developmental biology Developmental biology is the study of the process by which animals and plants grow and develop. Developmental biology also encompasses the biology of Regeneration (biology), regeneration, asexual reproduction, metamorphosis, and the growth and di ...
, ecology, morphology, and paleontology. Over 1.3 million insect species have been described, more than two-thirds of all known species. Some insect species date back to around 400 million years ago. They have many kinds of interactions with humans and other forms of life on Earth.


History

Entomology is rooted in nearly all human cultures from
prehistoric Prehistory, also known as pre-literary history, is the period of human history between the use of the first stone tools by hominins 3.3 million years ago and the beginning of recorded history with the invention of writing systems. The use of ...
times, primarily in the context of agriculture (especially biological control and
beekeeping Beekeeping (or apiculture) is the maintenance of bee colonies, commonly in man-made beehives. Honey bees in the genus '' Apis'' are the most-commonly-kept species but other honey-producing bees such as ''Melipona'' stingless bees are also kept. ...
). The natural philosopher Pliny the Elder, (23 - 79 AD) wrote a book on the kinds of Insects, while the scientist of Kufa,
Ibn al-A‘rābī Abū ʿAbd Allāh Muḥammad ibn Ziyād (), surnamed Ibn al-Aʿrābī () (ca. 760 – 846, Sāmarrā); a philologist, genealogist, and oral traditionist of Arabic tribal poetry. A grammarian of the school of al-Kūfah, who rivalled the grammari ...
(760 - 845 AD) wrote a book on flies, ''Kitāb al-Dabāb'' (). However scientific study in the modern sense began only relatively recently, in the 16th century. Ulisse Aldrovandi's ''De Animalibus Insectis'' (Of Insect Animals) was published in 1602. Microscopist
Jan Swammerdam Jan Swammerdam (February 12, 1637 – February 17, 1680) was a Dutch biologist and microscopist. His work on insects demonstrated that the various phases during the life of an insect—egg, larva, pupa, and adult—are different forms of the ...
published ''History of Insects'' correctly describing the reproductive organs of insects and
metamorphosis Metamorphosis is a biological process by which an animal physically develops including birth or hatching, involving a conspicuous and relatively abrupt change in the animal's body structure through cell growth and differentiation. Some inse ...
. In 1705, Maria Sibylla Merian published a book ''Metamorphosis Insectorum Surinamensium'' about the tropical insects of Dutch Surinam. Early entomological works associated with the naming and classification of species followed the practice of maintaining cabinets of curiosity, predominantly in Europe. This collecting fashion led to the formation of natural history societies, exhibitions of private collections, and journals for recording communications and the documentation of new species. Many of the collectors tended to be from the aristocracy and it spawned off a trade involving collectors around the world and traders. This has been called the "era of heroic entomology." William Kirby is widely considered as the father of entomology in England. In collaboration with William Spence, he published a definitive entomological encyclopedia, ''Introduction to Entomology'', regarded as the subject's foundational text. He also helped to found the Royal Entomological Society in London in 1833, one of the earliest such societies in the world; earlier antecedents, such as the Aurelian society date back to the 1740s. In the late 19th century, the growth of agriculture, and colonial trade, spawned off the "era of economic entomology" which created the professional entomologist associated with the rise of the university and training in the field of biology. Entomology developed rapidly in the 19th and 20th centuries, and was studied by large numbers of people, including such notable figures as Charles Darwin, Jean-Henri Fabre, Vladimir Nabokov, Karl von Frisch (winner of the 1973 Nobel Prize in Physiology or Medicine), and two-time
Pulitzer Prize The Pulitzer Prize () is an award for achievements in newspaper, magazine, online journalism, literature, and musical composition within the United States. It was established in 1917 by provisions in the will of Joseph Pulitzer, who had made h ...
winner E. O. Wilson. There has also been a history of people becoming entomologists through museum curation and research assistance, such as
Sophie Lutterlough Sophie Lutterlough (1910–2009) was an American entomologist. Lutterlough began working at the Smithsonian National Museum of Natural History (NMNH) as an elevator operator in the 1940s at a time when discriminatory hiring practices prevente ...
at the Smithsonian National Museum of Natural History. Insect identification is an increasingly common hobby, with
butterflies Butterflies are insects in the macrolepidopteran clade Rhopalocera from the Order (biology), order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The ...
and dragonflies being the most popular. Most insects can easily be recognized to order such as Hymenoptera (bees, wasps, and ants) or Coleoptera (beetles). However, identifying to genus or species is usually only possible through the use of identification keys and
monograph A monograph is a specialist work of writing (in contrast to reference works) or exhibition on a single subject or an aspect of a subject, often by a single author or artist, and usually on a scholarly subject. In library cataloging, ''monograph ...
s. Because the class Insecta contains a very large number of species (over 330,000 species of
beetles Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
alone) and the characteristics separating them are unfamiliar, and often subtle (or invisible without a microscope), this is often very difficult even for a specialist. This has led to the development of
automated species identification Automated species identification is a method of making the expertise of Taxonomy (biology), taxonomists available to ecologists, parataxonomy, parataxonomists and others via digital technology and artificial intelligence. Today, most automated ide ...
systems targeted on insects, for example, Daisy, ABIS, SPIDA and Draw-wing.


In pest control

In 1994, the Entomological Society of America launched a new
professional certification Professional certification, trade certification, or professional designation, often called simply ''certification'' or ''qualification'', is a designation earned by a person to assure qualification to perform a job or task. Not all certifications ...
program for the pest control industry called the Associate Certified Entomologist (ACE). To qualify as a "true entomologist" an individual would normally require an advanced degree, with most entomologists pursuing a PhD. While not true entomologists in the traditional sense, individuals who attain the ACE certification may be referred to as ACEs or Associate Certified Entomologists.


Subdisciplines

Many entomologists specialize in a single order or even a family of insects, and a number of these subspecialties are given their own informal names, typically (but not always) derived from the scientific name of the group: * Coleopterologybeetles. * Dipterologyflies. * Odonatologydragonflies and damselflies. * Hemipterologytrue bugs. * Isopterologytermites. * Lepidopterologymoths and
butterflies Butterflies are insects in the macrolepidopteran clade Rhopalocera from the Order (biology), order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The ...
. * Melittology (or ''Apiology'') –
bee Bees are winged insects closely related to wasps and ants, known for their roles in pollination and, in the case of the best-known bee species, the western honey bee, for producing honey. Bees are a monophyly, monophyletic lineage within the ...
s. * Myrmecologyants * Orthopterology
grasshopper Grasshoppers are a group of insects belonging to the suborder Caelifera. They are among what is possibly the most ancient living group of chewing herbivorous insects, dating back to the early Triassic around 250 million years ago. Grasshopp ...
s, crickets, etc. * Trichopterologycaddisflies. * Vespology – Social
wasps A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder. T ...
.


Entomologists


Organizations

Like other scientific specialties, entomologists have a number of local, national, and international organizations. There are also many organizations specializing in specific subareas. * Amateur Entomologists' Society *
Entomological Society of America The Entomological Society of America (ESA) was founded in 1889 and today has more than 7,000 members, including educators, extension personnel, consultants, students, researchers, and scientists from agricultural departments, health agencies, ...
*
Entomological Society of Canada The Entomological Society of Canada or Société d’Entomologie du Canada is one of Canada's most historic scientific societies. The society was founded in Toronto on April 16, 1863. The first Council was composed of President Henry Holmes Crof ...
*
Entomological Society of Japan The Entomological Society of Japan ( ja, 日本昆虫学会) was founded in 1917 for the purpose of improving and promoting entomology in Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It i ...
*
Entomologischer Verein Krefeld The Entomologischer Verein Krefeld (EVK) is an entomological society based in Krefeld, North Rhine-Westphalia, Germany. Founded in 1905, it keeps meticulous records and specimens of the area's insects, a collection curated since 1987 by entomologi ...

Entomological Society of India
*
International Union for the Study of Social Insects The International Union for the Study of Social Insects has at its purpose to promote and encourage the study of social insects and other social organisms in the broadest sense. Both research and the dissemination of knowledge about social insects ...
* Netherlands Entomological Society *
Royal Belgian Entomological Society The Royal Belgian Entomological Society (french: Société royale belge d'Entomologie; nl, Koninklijke Belgische Vereniging voor Entomologie) is a learned society based in Brussels. It is devoted to entomology, the study of insects. It was founded ...
* Royal Entomological Society of London * Russian Entomological Society *
Senckenberg Deutsches Entomologisches Institut The Senckenberg German Entomological Institute (german: Senckenberg Deutsches Entomologisches Institut, SDEI or DEI) is a German entomological research institute devoted to the study of insects. Founded in 1886, the institute has an extraordina ...
* Société entomologique de France
Australian Entomological Society

Entomological Society of New Zealand


Research collection

Here is a list of selected very large insect collections, housed in museums, universities, or research institutes.


Asia

*
Zoological Survey of India The Zoological Survey of India (ZSI), founded on 1 July 1916 by Government of India Ministry of Environment, Forest and Climate Change, as premier Indian organisation in zoological research and studies to promote the survey, exploration and r ...
*
Insect Museum, Tamil Nadu Agricultural University, Coimbatore, Tamil Nadu, India Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (head, thorax and abdomen), three pairs of j ...
* National Pusa Collection, Division of Entomology, Indian Agricultural Research Institute, New Delhi, India *
Pakistan Museum of Natural History Pakistan Museum of Natural History (PMNH), ( ur, ) established in 1976, is a public natural history museum situated in Islamabad, the federal capital of Pakistan. It has exhibits and galleries which display and provide information about the ecol ...
Garden Avenue, Shakarparian, Islamabad, Pakistan * Museum Zoologicum Bogoriense, Indonesia


Africa

* Natal Museum,
Pietermaritzburg Pietermaritzburg (; Zulu: umGungundlovu) is the capital and second-largest city in the province of KwaZulu-Natal, South Africa. It was founded in 1838 and is currently governed by the Msunduzi Local Municipality. Its Zulu name umGungundlovu ...
, South Africa


Australasia

*
Lincoln University Entomology Research Collection The Lincoln University Entomology Research Collection is a collection of approximately 500,000 insect, spider, and other arthropod specimens housed in Lincoln University (New Zealand), Lincoln University, New Zealand. One of New Zealand's larges ...
, Lincoln, New Zealand *
Museum of New Zealand Te Papa Tongarewa The Museum of New Zealand Te Papa Tongarewa is New Zealand's national museum and is located in Wellington. ''Te Papa Tongarewa'' translates literally to "container of treasures" or in full "container of treasured things and people that spring fr ...
, Wellington, New Zealand * New Zealand Arthropod Collection, Landcare Research Manaaki Whenua, Auckland, New Zealand


Europe

* Bavarian State Collection of Zoology,
Zoologische Staatssammlung München The Bavarian State Collection of Zoology (german: Zoologische Staatssammlung München) or ZSM is a major German research institution for zoological systematics in Munich. It has over 20 million zoological specimens. It is one of the largest natur ...
* Museu de Ciències Naturals de Barcelona, Barcelona, Spain * Muséum national d'histoire naturelle, Paris, France * Museum für Naturkunde, Berlin, Germany *
Kelvingrove Art Gallery Kelvingrove Art Gallery and Museum is a museum and art gallery in Glasgow, Scotland. It reopened in 2006 after a three-year refurbishment and since then has been one of Scotland's most popular visitor attractions. The museum has 22 galleries, h ...
, Glasgow, Scotland * Natural History Museum, Budapest Hungarian Natural History Museum * Natural History Museum, Geneva *
Natural History Museum, Leiden Naturalis Biodiversity Center ( nl, Nederlands Centrum voor Biodiversiteit Naturalis) is a national museum of natural history and a research center on biodiversity in Leiden, Netherlands. It was named the European Museum of the Year 2021. Altho ...
, the Netherlands * Natural History Museum, London, United Kingdom *
Natural History Museum, Oslo The Natural History Museum at the University of Oslo ( no, Naturhistorisk museum, NHM) is Norway's oldest and largest museum of natural history. It is situated in the neighborhood of Tøyen in Oslo, Norway. It traces its roots to the Universit ...
Norway * Natural History Museum, St. Petersburg Zoological Collection of the Russian Academy of Science * Naturhistorisches Museum, Vienna, Austria * Oxford University Museum of Natural History, Oxford * Royal Museum for Central Africa, Brussels, Belgium *
Swedish Museum of Natural History The Swedish Museum of Natural History ( sv, Naturhistoriska riksmuseet, literally, the National Museum of Natural History), in Stockholm, is one of two major museums of natural history in Sweden, the other one being located in Gothenburg. The ...
,
Stockholm Stockholm () is the Capital city, capital and List of urban areas in Sweden by population, largest city of Sweden as well as the List of urban areas in the Nordic countries, largest urban area in Scandinavia. Approximately 980,000 people liv ...
,
Sweden Sweden, formally the Kingdom of Sweden,The United Nations Group of Experts on Geographical Names states that the country's formal name is the Kingdom of SwedenUNGEGN World Geographical Names, Sweden./ref> is a Nordic country located on ...
*
World Museum Liverpool World Museum is a large museum in Liverpool, England which has extensive collections covering archaeology, ethnology and the natural and physical sciences. Special attractions include the Natural History Centre and a planetarium. Entry to the ...
, the ''Bug House''


United States

* Academy of Natural Sciences of Philadelphia *
American Museum of Natural History The American Museum of Natural History (abbreviated as AMNH) is a natural history museum on the Upper West Side of Manhattan in New York City. In Theodore Roosevelt Park, across the street from Central Park, the museum complex comprises 26 inter ...
, New York City * Auburn University Museum of Natural History, Auburn, Alabama * Audubon Insectarium, New Orleans *
Bohart Museum of Entomology The Bohart Museum of Entomology was founded in 1946 on the campus of the University of California, Davis. The museum is currently the seventh largest insect collection in North America with more than seven million specimens of terrestrial and f ...
, Davis, California * California Academy of Sciences, San Francisco * Carnegie Museum of Natural History, Pittsburgh * Cleveland Museum of Natural History, Cleveland * Entomology Research Museum, University of California, Riverside *
Essig Museum of Entomology The Essig Museum of Entomology is a research museum dedicated to terrestrial arthropods, located at the University of California at Berkeley. It contains perhaps the world's largest collection of California insects and has been ranked among the top ...
, Berkeley, California *
Field Museum of Natural History The Field Museum of Natural History (FMNH), also known as The Field Museum, is a natural history museum in Chicago, Illinois, and is one of the largest such museums in the world. The museum is popular for the size and quality of its educational ...
, Chicago *
Florida Museum of Natural History The Florida Museum of Natural History (FLMNH) is Florida's official state-sponsored and chartered natural-history museum. Its main facilities are located at 3215 Hull Road on the campus of the University of Florida in Gainesville. The main pub ...
, University of Florida,
Gainesville, Florida Gainesville is the county seat of Alachua County, Florida, Alachua County, Florida, and the largest city in North Central Florida, with a population of 141,085 in 2020. It is the principal city of the Gainesville metropolitan area, Florida, Gaine ...
* Illinois Natural History Survey, Champaign, Illinois *
J. Gordon Edwards Museum ''J. The Jewish News of Northern California'', formerly known as ''Jweekly'', is a weekly print newspaper in Northern California, with its online edition updated daily. It is owned and operated by San Francisco Jewish Community Publications In ...
, San Jose, California * Museum of Comparative Zoology, Cambridge, Massachusetts * Natural History Museum of Los Angeles County, Los Angeles * National Museum of Natural History, Washington, D.C. *
New Mexico State University Arthropod Museum New is an adjective referring to something recently made, discovered, or created. New or NEW may refer to: Music * New, singer of K-pop group The Boyz Albums and EPs * ''New'' (album), by Paul McCartney, 2013 * ''New'' (EP), by Regurgitator, ...
*
North Carolina State University Insect Museum The NCSU Insect Museum is the center for research and training in insect systematics and biodiversity informatics at North Carolina State University. The Museum's collections hold more than 1.5 million specimens, with major emphases on the insec ...
, Raleigh, North Carolina * Peabody Museum of Natural History, New Haven, Connecticut * San Diego Natural History Museum, San Diego, California *
The National Museum of Play The Strong National Museum of Play (known as just The Strong Museum or simply the Strong) is part of The Strong in Rochester, New York, United States. Established in 1969 and based initially on the personal collection of Rochester native Margaret ...
, Rochester, N.Y. * Texas A&M University, College Station, Texas * University of Minnesota, St. Paul campus (
UMSP The University of Minnesota, formally the University of Minnesota, Twin Cities, (UMN Twin Cities, the U of M, or Minnesota) is a public land-grant research university in the Twin Cities of Minneapolis and Saint Paul, Minnesota, United States. T ...
), Minnesota * University of Kansas Natural History Museum,
Lawrence, Kansas Lawrence is the county seat of Douglas County, Kansas, Douglas County, Kansas, United States, and the sixth-largest city in the state. It is in the northeastern sector of the state, astride Interstate 70, between the Kansas River, Kansas and Waka ...
* University of Nebraska State Museum, Lincoln, Nebraska * University of Missouri Enns Entomology Museum, University of Missouri,
Columbia, Missouri Columbia is a city in the U.S. state of Missouri. It is the county seat of Boone County and home to the University of Missouri. Founded in 1821, it is the principal city of the five-county Columbia metropolitan area. It is Missouri's fourth ...


Canada

*
Canadian Museum of Nature The Canadian Museum of Nature (french: Musée canadien de la nature; CMN) is a national natural history museum based in Canada's National Capital Region. The museum's exhibitions and public programs are housed in the Victoria Memorial Museum Bui ...
,
Ottawa Ottawa (, ; Canadian French: ) is the capital city of Canada. It is located at the confluence of the Ottawa River and the Rideau River in the southern portion of the province of Ontario. Ottawa borders Gatineau, Quebec, and forms the core ...
, Ontario * Canadian National Collection of Insects, Arachnids and Nematodes,
Ottawa Ottawa (, ; Canadian French: ) is the capital city of Canada. It is located at the confluence of the Ottawa River and the Rideau River in the southern portion of the province of Ontario. Ottawa borders Gatineau, Quebec, and forms the core ...
, Ontario * E.H. Strickland Entomological Museum, University of Alberta, Edmonton, Alberta * Lyman Entomological Museum, Macdonald Campus of McGill University, Sainte-Anne-de-Bellevue, Quebec * Montreal Insectarium, Montreal, Quebec *
Newfoundland Insectarium The Newfoundland Insectarium is a museum insectarium located in Reidville, Newfoundland and Labrador, Canada. The main display features a wide variety of mounted insects from around the world, organized by geographical region. One exhibit cover ...
,
Reidville, Newfoundland and Labrador Reidville is a village located north east of Deer Lake. A post office was established in 1967 and the first Postmisstress was Dorothy Barrett. Demographics In the 2021 Census of Population conducted by Statistics Canada, Reidville had a popul ...
* Royal Alberta Museum, Edmonton, Alberta *
Royal Ontario Museum The Royal Ontario Museum (ROM) is a museum of art, world culture and natural history in Toronto, Ontario, Canada. It is one of the largest museums in North America and the largest in Canada. It attracts more than one million visitors every year ...
, Toronto, Ontario * University of Guelph Insect Collection, Guelph, Ontario * Victoria Bug Zoo, Victoria, British Columbia * J. B. Wallis / R. E. Roughley Museum of Entomology, Winnipeg, Manitoba


See also

* Arachnology * Carcinology * Cultural entomology *
Ethnoentomology Human interactions with insects include both a wide variety of uses, whether practical such as for food, textiles, and dyestuffs, or symbolic, as in art, music, and literature, and negative interactions including serious damage to crops and exten ...
* Forensic entomology *
Forensic entomologist Forensic entomology is the scientific study of the colonization of a dead body by arthropods. This includes the study of insect types commonly associated with cadavers, their respective life cycles, their ecological presences in a given environme ...
*
Forensic entomology and the law Forensic entomology deals with the collection of arthropodic evidence and its application, and through a series of tests and previously set of rules, general admissibility of said evidence is determined. Forensic entomology may come into play in a ...
* Insect thermoregulation *
Insects on stamps Many countries have featured insects on stamps. Insect related topics such as the mosquito eradication (anti malaria) programme of the 1960s as well as graphic designs based on insects have also appeared. Many stamps also feature butterflies. Ins ...
* List of entomological journals * Medical entomology * Myriapodology *
Timeline of entomology – 1800–1850 19th century 1800 – an arbitrary date but it was around this time that systematists began to specialise. There remained entomological polyhistors – those who continued to work on the insect fauna as a whole. From the beginning of the cent ...
*
Timeline of entomology – 1850–1900 1850 *Edmond de Sélys Longchamps . 6:1–408. *Victor Ivanovitsch Motschulsky . I. ''Insecta Carabica''. Russian beetles, Carabidae, Moscow: Gautier, published. 1851 *Johann Fischer von Waldheim and Eduard Friedrich Eversmann publish (vol.5 ...
*
Timeline of entomology since 1900 ;1900 * Walter Reed, a United States Army major, was appointed president of a board "to study infectious diseases in Cuba paying particular attention to yellow fever." He concurred with Carlos Finlay in identifying mosquitoes as the agent. * I ...


References


Further reading

* Chiang, H.C. and G. C. Jahn 1996. Entomology in the Cambodia-IRRI-Australia Project. (in Chinese) ''Chinese Entomol. Soc. Newsltr.'' (Taiwan) 3: 9–11. * Davidson, E. 2006. ''Big Fleas Have Little Fleas: How Discoveries of Invertebrate Diseases Are Advancing Modern Science'' University of Arizona Press, Tucson, 208 pages, . * Cedric Gillot: '' Entomology''. Second Edition, Plenum Press, New York, NY / London 1995, . * Triplehorn, Charles A. and Norman F. Johnson (2005-05-19). Borror and DeLong's Introduction to the Study of Insects, 7th edition, Thomas Brooks/Cole. . — a classic textbook in North America. * * Capinera, JL (editor). 2008. ''Encyclopedia of Entomology'', 2nd Edition. Springer.


External links

{{Authority control Subfields of arthropodology