''Enophrys diceraus'', the antlered sculpin, is a species of marine

ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or hor ...

belonging to the

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Cottidae The Cottidae are a family of fish in the superfamily Cottoidea, the sculpins. It is the largest sculpin family, with about 275 species in 70 genera.Kane, E. A. and T. E. Higham. (2012)Life in the flow lane: differences in pectoral fin morphology ...

, the typical sculpins. This species occurs in the northern Pacific Ocean.

Taxonomy

''Enophrys diceraus'' was first formally described as ''Cottus diceraus'' in 1787 by the German

zoologist Zoology ()The pronunciation of zoology as is usually regarded as nonstandard, though it is not uncommon. is the branch of biology that studies the Animal, animal kingdom, including the anatomy, structure, embryology, evolution, Biological clas ...

Peter Simon Pallas Peter Simon Pallas Fellow of the Royal Society, FRS FRSE (22 September 1741 – 8 September 1811) was a Prussian zoologist and botanist who worked in Russia between 1767 and 1810. Life and work Peter Simon Pallas was born in Berlin, the son ...

with its type locality given as Kamchatka. In 1829 the French zoologist

Georges Cuvier Jean Léopold Nicolas Frédéric, Baron Cuvier (; 23 August 1769 – 13 May 1832), known as Georges Cuvier, was a French natural history, naturalist and zoology, zoologist, sometimes referred to as the "founding father of paleontology". Cuvier ...

described ''Cottus claviger'', also from Kamchatka, and in 1839 the English zoologist

William John Swainson William John Swainson FLS, FRS (8 October 1789 – 6 December 1855), was an English ornithologist, malacologist, conchologist, entomologist and artist. Life Swainson was born in Dover Place, St Mary Newington, London, the eldest son of ...

proposed a new

monospecific genus In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...

for Cuvier's species which he called ''

Enophrys ''Enophrys'' is a genus of marine ray-finned fishes belonging to the family Cottidae, the typical sculpins. These fishes are found in the northern and eastern Pacific Ocean. Taxonomy ''Enophrys'' was first proposed as a monospecific genus in 183 ...

''. Later Cuvier's ''C. claviger'' was found to be a

synonym A synonym is a word, morpheme, or phrase that means exactly or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are all ...

of Pallas's ''C, diceraus'' but it is the

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

of the genus ''Enophrys''. The

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

''diceraus'' means "two-horned" a reference to the preopercular spines, one on each side of the head.

Description

''Enophrys diceraus'' has a long straight spine on the upper cheek, the preopercular spine which reaches past the operculum and has between 2 and 8 ancillary spines, or spinules. The lacrimal bone has a bifurcated flap which extends down over the

maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The t ...

. There are 2

dorsal fin A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through conv ...

s which are supported by between 7 or 8 spines and between 12 and 15 soft rays while the

anal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...

has 10 or 13 soft rays. There are 2 well developed bony ridges on the top of the head. The large

lateral line The lateral line, also called the lateral line organ (LLO), is a system of sensory organs found in fish, used to detect movement, vibration, and pressure gradients in the surrounding water. The sensory ability is achieved via modified epithelial ...

scales are plate like and have 1 or 2 keels and a rough rera edge. This species reaches a maximum published

total length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish ...

of .

Distribution and habitat

''Enophrys diceraus'' is found in the subarctic part of the Pacific Ocean from the northern

Sea of Japan The Sea of Japan is the marginal sea between the Japanese archipelago, Sakhalin, the Korean Peninsula, and the mainland of the Russian Far East. The Japanese archipelago separates the sea from the Pacific Ocean. Like the Mediterranean Sea, it h ...

Point Hope Point Hope ( ik, Tikiġaq, ) is a city in North Slope Borough, Alaska, United States. At the 2010 census the population was 674, down from 757 in 2000. In the 2020 Census, population rose to 830. Like many isolated communities in Alaska, the c ...

Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...

and in the

Chukchi Sea Chukchi Sea ( rus, Чуко́тское мо́ре, r=Chukotskoye more, p=tɕʊˈkotskəjə ˈmorʲɪ), sometimes referred to as the Chuuk Sea, Chukotsk Sea or the Sea of Chukotsk, is a marginal sea of the Arctic Ocean. It is bounded on the west b ...

Amchitka Island Amchitka (; ale, Amchixtax̂; russian: Амчитка) is a volcanic, tectonically unstable and uninhabited island in the Rat Islands group of the Aleutian Islands in southwest Alaska. It is part of the Alaska Maritime National Wildlife Refu ...

in the

Aleutians The Aleutian Islands (; ; ale, Unangam Tanangin,”Land of the Aleuts", possibly from Chukchi ''aliat'', "island"), also called the Aleut Islands or Aleutic Islands and known before 1867 as the Catherine Archipelago, are a chain of 14 large vo ...

. It is a

benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning "t ...

species of coastal waters dound at depths between over substrates of sand, mud and rock.

Biology

''Enophrys diceraus'' eats benthic invertebrates, especially crustaceans and molluscs, although adults eat

brittlestar Brittlestar is the stage name of Stewart Reynolds, a Stratford, Ontario based comedian, writer, communications consultant, and online television show host. His campaign for KFC Canada was the world's most watched branded video on Facebook in th ...

s too, The adults move into shallow water to spawn, the juvenile settle at lengths between and they become sexually mature at . They form aggregations for feeding and spawning in the south of their range in

Peter the Great Bay The Peter the Great Gulf (Russian: Залив Петра Великого) is a gulf on the southern coast of Primorsky Krai, Russia, and the largest gulf of the Sea of Japan. The gulf extends for from the Russian-North Korean border at the mout ...

and in central parts along the mainland Asian coast and off the southern part of

Sakhalin Sakhalin ( rus, Сахали́н, r=Sakhalín, p=səxɐˈlʲin; ja, 樺太 ''Karafuto''; zh, c=, p=Kùyèdǎo, s=库页岛, t=庫頁島; Manchu: ᠰᠠᡥᠠᠯᡳᠶᠠᠨ, ''Sahaliyan''; Orok: Бугата на̄, ''Bugata nā''; Nivkh: ...

. These do not occur in the northern part of their range.

References

{{Taxonbar, from=Q2440775 Fish described in 1787 Taxa named by Peter Simon Pallas diceraus