''Enhydra'' is a genus of mustelid that contains the
sea otter
The sea otter (''Enhydra lutris'') is a marine mammal native to the coasts of the northern and eastern North Pacific Ocean. Adult sea otters typically weigh between , making them the heaviest members of the weasel family, but among the small ...
and two extinct relatives. It is the only extant genus of the bunodont otters group, referring to otters with non-blade
carnassial
Carnassials are paired upper and lower teeth modified in such a way as to allow enlarged and often self-sharpening edges to pass by each other in a shearing manner. This adaptation is found in carnivorans, where the carnassials are the modified f ...
s with rounded cusps.
Sea otters probably diverged from other otters during the Pliocene, approximately 5 mya. They probably arose from the closely related ''
Enhydritherium
''Enhydritherium terraenovae'' is an extinct marine otter endemic to North America that lived during the Miocene through Pliocene epochs from ~9.1–4.9 Ma. ( AEO), existing for approximately .
The ancestral lineage of ''Enhydritherium terraen ...
'', a bunodont otter endemic to North America during the late
Miocene
The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
and early
Pliocene
The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58
''
Enhydra reevei
''Enhydra'' is a genus of mustelid that contains the sea otter and two extinct relatives. It is the only extant genus of the bunodont otters group, referring to otters with non-blade carnassials with rounded cusps.
Sea otters probably diverged f ...
'', the oldest known species, has its origins in the Atlantic, suggesting this may have been where sea otters originated. Fossil evidence indicates the ''Enhydra'' lineage became isolated in the
North Pacific
The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continen ...
approximately 2 million years ago, giving rise to the now-extinct ''
Enhydra macrodonta
''Enhydra macrodonta'', the large-toothed sea otter, is an extinct mustelid known from the middle Pleistocene in California
California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2million resid ...