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Engis 2 refers to part of an assemblage, discovered in 1829 by Dutch physician and naturalist
Philippe-Charles Schmerling Philippe-Charles or Philip Carel Schmerling (2 March 1791 Delft Р7 November 1836, Li̬ge) was a Dutch/Belgian prehistorian, pioneer in paleontology, and geologist. He is often considered the founder of paleontology. In 1829 he discovered ...
in the lower of the
Schmerling Caves The Schmerling Caves (also known as Grottes d'Engis, meaning Engis Caves) are a group of caves located in Wallonia on the right bank of the stream called the Awirs, near the village of Awirs in Flémalle, Belgium. The caves are notable for their p ...
. The pieces that make up Engis 2 are a partially preserved calvaria (cranium) and associated fragments of an upper and a lower jaw, a
maxillary bone The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The tw ...
and an upper
incisor Incisors (from Latin ''incidere'', "to cut") are the front teeth present in most mammals. They are located in the premaxilla above and on the mandible below. Humans have a total of eight (two on each side, top and bottom). Opossums have 18, whe ...
tooth of a two to three year old
Neanderthal Neanderthals (, also ''Homo neanderthalensis'' and erroneously ''Homo sapiens neanderthalensis''), also written as Neandertals, are an extinct species or subspecies of archaic humans who lived in Eurasia until about 40,000 years ago. While th ...
child. The
Schmerling Caves The Schmerling Caves (also known as Grottes d'Engis, meaning Engis Caves) are a group of caves located in Wallonia on the right bank of the stream called the Awirs, near the village of Awirs in Flémalle, Belgium. The caves are notable for their p ...
are situated just north of the
Belgian Belgian may refer to: * Something of, or related to, Belgium * Belgians, people from Belgium or of Belgian descent * Languages of Belgium, languages spoken in Belgium, such as Dutch, French, and German *Ancient Belgian language, an extinct languag ...
municipality A municipality is usually a single administrative division having corporate status and powers of self-government or jurisdiction as granted by national and regional laws to which it is subordinate. The term ''municipality'' may also mean the go ...
Engis Engis (; wa, Indji) is a municipality of Wallonia located in the province of Liège, Belgium. On 1 January 2006 Engis had a total population of 5,686. The total area is 27.74 km² which gives a population density of 205 inhabitants per km ...
, whence the name of this group. In 1833 Schmerling described and publicized the find, which included animal bones and stone tools. Recognizing their old age, he associated them with the "Ethiopian Type" of the diluvial period. Although it was not recognized as such until 1936, the publication represents the first scientific description of a
Neanderthal Neanderthals (, also ''Homo neanderthalensis'' and erroneously ''Homo sapiens neanderthalensis''), also written as Neandertals, are an extinct species or subspecies of archaic humans who lived in Eurasia until about 40,000 years ago. While th ...
fossil.


Early Misclassification

Originally misclassified as "modern", the fossil received little attention after its publication in the 19th century as it was compared to Engis 1 - the very good and almost perfectly preserved skull of an adult ''
Homo sapiens Humans (''Homo sapiens'') are the most abundant and widespread species of primate, characterized by bipedalism and exceptional cognitive skills due to a large and complex brain. This has enabled the development of advanced tools, culture, ...
''. In 1758
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
had published the 10th edition of his work
Systema Naturae ' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial nomen ...
in which ''
Homo sapiens Humans (''Homo sapiens'') are the most abundant and widespread species of primate, characterized by bipedalism and exceptional cognitive skills due to a large and complex brain. This has enabled the development of advanced tools, culture, ...
'' as a species name was introduced to the public, yet without a thorough diagnosis and without a precise description of the species-specific characteristics. As a result, any criteria by which a fossil of the
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
''Homo sapiens'' could be classified into and distinguished from the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''
Homo ''Homo'' () is the genus that emerged in the (otherwise extinct) genus ''Australopithecus'' that encompasses the extant species ''Homo sapiens'' ( modern humans), plus several extinct species classified as either ancestral to or closely relate ...
'' did not exist in the early 19th century. Even
Thomas Henry Huxley Thomas Henry Huxley (4 May 1825 – 29 June 1895) was an English biologist and anthropologist specialising in comparative anatomy. He has become known as "Darwin's Bulldog" for his advocacy of Charles Darwin's theory of evolution. The storie ...
, a supporter of Darwin's theory of
evolution Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variation ...
, saw in the 1863 findings of the Engis cave a "man of low degree of civilization" and also interpreted the
Neandertal 1 Feldhofer 1 or Neanderthal 1 is the scientific name of the 40,000-year-old Type (biology), type specimen fossil of the species Neanderthal, ''Homo neanderthalensis'', found in August 1856 in a Germany, German cave, the Kleine Feldhofer Grotte in ...
fossils of the
Kleine Feldhofer Grotte Kleine Feldhofer Grotte was a karstic limestone cave and a paleoanthropologic site in the Neander Valley in western Germany. In August 1856, the Neanderthal type specimen was unearthed from the cave. Miners uncovered a skull cap and a number of ...
unearthed in 1856 as belonging within the range of variations of modern man. Additionally, the skull of an infant Neanderthal and an equally old child of anatomically modern humans are of far greater resemblance than their respective adult skulls. The vast majority of the
anthropologists An anthropologist is a person engaged in the practice of anthropology. Anthropology is the study of aspects of humans within past and present societies. Social anthropology, cultural anthropology and philosophical anthropology study the norms and v ...
of the 19th and early 20th century considered all hominid fossils as belonging to representatives of early "races" of modern man. Hence it was incorrectly believed that the modern man's skull Engis 1 must be related to the child's skull Engis 2.


Reclassification and Age

Two radiocarbon dates are available for Engis 2. However, the earlier result of 26,820 ± 340 radiocarbon years before present (BP) was considered to be too young by the authors and likely to be a result of contamination, so has been discarded. The more accurate date is 30,460 ± 210 radiocarbon years BP, which corresponds to 34,590–36,110 years BP after calibration. The assignment of Engis 2 to ''
Homo neanderthalensis Neanderthals (, also ''Homo neanderthalensis'' and erroneously ''Homo sapiens neanderthalensis''), also written as Neandertals, are an Extinction, extinct species or subspecies of archaic humans who lived in Eurasia until about 40,000 years ag ...
'' and Engis 1 to ''Homo sapiens'' was primarily based on anatomical and chronological comparisons as Engis 2 was recovered in the context of
Neanderthal Neanderthals (, also ''Homo neanderthalensis'' and erroneously ''Homo sapiens neanderthalensis''), also written as Neandertals, are an extinct species or subspecies of archaic humans who lived in Eurasia until about 40,000 years ago. While th ...
Mousterian The Mousterian (or Mode III) is an archaeological industry of stone tools, associated primarily with the Neanderthals in Europe, and to the earliest anatomically modern humans in North Africa and West Asia. The Mousterian largely defines the latt ...
artefacts.


Condition of the bones and associated assemblages

In 1986 cut marks were found on the top of the skull of Engis 2, which were later identified as to be preparation damage " formed during restoration of the vault, moulding striae formed when mold part lines were incised into the fossil and profiling striae formed when craniograms were made with sharp steel instrument tips." The findings are preserved at the ''Collections de Paléontologie Animale et Humaine'' of the
University of Liège The University of Liège (french: Université de Liège), or ULiège, is a major public university of the French Community of Belgium based in Liège, Wallonia, Belgium. Its official language is French. As of 2020, ULiège is ranked in the 301 ...
. The bone fragments called Engis 3 have gone missing. The evolutionary origin of an
ulna The ulna (''pl''. ulnae or ulnas) is a long bone found in the forearm that stretches from the elbow to the smallest finger, and when in anatomical position, is found on the medial side of the forearm. That is, the ulna is on the same side of t ...
(forearm bone) fragment called Engis 4 discovered in 1872 is unclear; it has to date not been associated with a specific
taxon In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...
.


See also

*
List of fossil sites This list of fossil sites is a worldwide list of localities known well for the presence of fossils. Some entries in this list are notable for a single, unique find, while others are notable for the large number of fossils found there. Many of t ...
''(with link directory)'' *
List of human evolution fossils The following tables give an overview of notable finds of Hominini, hominin fossils and Skeleton, remains relating to human evolution, beginning with the formation of the tribe Hominini (the divergence of the Chimpanzee–human last common ancest ...
''(with images)'' *
Neanderthal Neanderthals (, also ''Homo neanderthalensis'' and erroneously ''Homo sapiens neanderthalensis''), also written as Neandertals, are an extinct species or subspecies of archaic humans who lived in Eurasia until about 40,000 years ago. While th ...
*
Neanderthal 1 Feldhofer 1 or Neanderthal 1 is the scientific name of the 40,000-year-old type specimen fossil of the species ''Homo neanderthalensis'', found in August 1856 in a German cave, the Kleine Feldhofer Grotte in the Neandertal valley, east of Düs ...
*
Neanderthals of Gibraltar The Neanderthals in Gibraltar were among the first to be discovered by modern scientists and have been among the most well studied of their species according to a number of extinction studies which emphasize regional differences, usually claiming ...


References


External links

*
Science/AAAS - Background - The Neandertal Genome

Engis 2 - The Smithsonian Institution - Human Origins Program
{{Homo neanderthalensis, state=expanded Neanderthal fossils 1829 in the Southern Netherlands 1829 archaeological discoveries Archaeology of Belgium