''Endothenia quadrimaculana'' is a

moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...

of the family Tortricidae. It is found from northern and central

Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...

Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...

and south-eastern

Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...

Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million, ...

and

China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...

. Subspecies ''nubilana'' is found in

North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...

. The

wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingspan of ...

is 18–22 mm.The ground colour of the forewings is light brown with small dark brown spots along the costal edge, a square brown spot in the middle of the wing and an inward-curved brown cross-band at the tip. The hindwings are light brown. Dissection of the genitalia is necessary to determine ''Endothenia'' species with certainty. In France and Switzerland, there are two generations per year. Adults are on wing in May and June and again in August and September. The larvae feed on '' Mentha spicata'', ''

Mentha arvensis ''Mentha arvensis'', the corn mint, field mint, or wild mint, is a species of flowering plant in the mint family Lamiaceae. It has a circumboreal distribution, being native to the temperate regions of Europe and western and central Asia, east to ...

'', ''

Lamium album ''Lamium album'', commonly called white nettle or white dead-nettle, is a flowering plant in the family Lamiaceae. It is native throughout Europe and Asia, growing in a variety of habitats from open grassland to woodland, generally on moist, fert ...

'', ''

Stachys palustris ''Stachys palustris'', commonly known as marsh woundwort, clown's woundwort, clown's heal-all, marsh hedgenettle, or hedge-nettle,Dickinson, T.; Metsger, D.; Bull, J.; & Dickinson, R. (2004) ROM Field Guide to Wildflowers of Ontario. Toronto:Roya ...

'', ''

Stachys arvensis ''Stachys arvensis'' is a species of flowering plant in the mint family known by the common names field woundwort and staggerweed. It is native to Europe, Western Asia, and North Africa. It is known on other continents as an introduced species an ...

'', ''

Stachys recta ''Stachys recta'', commonly known as stiff hedgenettle or perennial yellow-woundwort, is herbaceous perennial plant of the family Lamiaceae. Etymology The generic epithet is derived from the Greek word σταχυς (''stachys''), meaning "an ear ...

'' and ''

Symphytum officinale ''Symphytum officinale'' is a perennial flowering plant in the family Boraginaceae. Along with thirty four other species of ''Symphytum'', it is known as comfrey. To differentiate it from other members of the genus ''Symphytum'', this species is ...

''. In France also on ''

Stachys affinis ''Stachys affinis'', commonly called crosne, Chinese artichoke, Japanese artichoke, knotroot, or artichoke betony, is a perennial herbaceous plant of the family Lamiaceae, originating from China. Its rhizome is a root vegetable that can be e ...

'', which is imported from Japan. The larvae attack and damage the rhizomes of '' Stachys'' species, which will spoil easily and have to be eliminated before being sold. Only the larvae of the second generation cause economically important damage to cultivated Stachys, those of the early generation tunnel in the roots and the runners.

Subspecies

*''Endothenia quadrimaculana quadrimaculana'' (Eurasia) *''Endothenia quadrimaculana nubilana'' (North America)

External links

Eurasian TortricidaeUKmoths
Endotheniini Moths described in 1963 Moths of Japan Moths of Europe Moths of Asia {{Olethreutinae-stub