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''Endemoconus'' is
subgenus In biology, a subgenus (plural: subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between t ...
of
sea snail Sea snail is a common name for slow-moving marine gastropod molluscs, usually with visible external shells, such as whelk or abalone. They share the taxonomic class Gastropoda with slugs, which are distinguished from snails primarily by the ...
s,
marine Marine is an adjective meaning of or pertaining to the sea or ocean. Marine or marines may refer to: Ocean * Maritime (disambiguation) * Marine art * Marine biology * Marine debris * Marine habitats * Marine life * Marine pollution Military * ...
gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. T ...
mollusk Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is e ...
s in the genus ''
Conasprella ''Conasprella'' is a genus of sea snails, marine gastropod mollusks in the family Conidae, the cone snails and their allies.Bouchet, P. (2015). Conasprella. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia ...
'',
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Conidae Conidae, with the current common name of "cone snails", is a taxonomic family (previously subfamily) of predatory sea snails, marine gastropod molluscs in the superfamily Conoidea. The 2014 classification of the superfamily Conoidea, groups onl ...
, the
cone snails A cone is a three-dimensional geometric shape that tapers smoothly from a flat base (frequently, though not necessarily, circular) to a point called the apex or vertex. A cone is formed by a set of line segments, half-lines, or lines co ...
and their allies.Bouchet, P. (2015). Endemoconus Iredale, 1931. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=430148 on 2015-03-17 In the new classification of the family Conidae by Puillandre N., Duda T.F., Meyer C., Olivera B.M. & Bouchet P. (2015), ''Endemoconus'' has become a subgenus of ''Conasprella'': ''Conasprella (Endemoconus)'' Tucker & Tenorio, 2009 represented as ''
Conasprella ''Conasprella'' is a genus of sea snails, marine gastropod mollusks in the family Conidae, the cone snails and their allies.Bouchet, P. (2015). Conasprella. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia ...
'' Thiele, 1929. Three species have become synonyms of a ''Conus'' species. Puillandre N., Duda T.F., Meyer C., Olivera B.M. & Bouchet P. (2015). ''One, four or 100 genera? A new classification of the cone snails''. Journal of Molluscan Studies. 81: 1-23
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Distinguishing characteristics

The Tucker & Tenorio 2009 taxonomy distinguishes ''Endemoconus'' from ''Conus'' in the following ways:Tucker J.K. & Tenorio M.J. (2009), Systematic Classification of Recent and Fossil Conoidean Gastropods, ConchBooks, Hankenheim, Germany, 295 pp. * Genus ''Conus'' ''
sensu stricto ''Sensu'' is a Latin word meaning "in the sense of". It is used in a number of fields including biology, geology, linguistics, semiotics, and law. Commonly it refers to how strictly or loosely an expression is used in describing any particular co ...
'' Linnaeus, 1758 :: Shell characters (living and fossil species) :::The basic shell shape is conical to elongated conical, has a deep anal notch on the shoulder, a smooth
periostracum The periostracum ( ) is a thin, organic coating (or "skin") that is the outermost layer of the shell of many shelled animals, including molluscs and brachiopods. Among molluscs, it is primarily seen in snails and clams, i.e. in gastropods and ...
and a small operculum. The shoulder of the shell is usually nodulose and the
protoconch A protoconch (meaning first or earliest or original shell) is an embryonic or larval shell which occurs in some classes of molluscs, e.g., the initial chamber of an ammonite or the larval shell of a gastropod. In older texts it is also called ...
is usually multispiral. Markings often include the presence of tents except for black or white color variants, with the absence of spiral lines of minute tents and textile bars. ::Radular tooth (not known for fossil species) :::The
radula The radula (, ; plural radulae or radulas) is an anatomical structure used by molluscs for feeding, sometimes compared to a tongue. It is a minutely toothed, chitinous ribbon, which is typically used for scraping or cutting food before the food ...
has an elongated anterior section with serrations and a large exposed terminating cusp, a non-obvious waist, blade is either small or absent and has a short barb, and lacks a basal spur. ::Geographical distribution :::These species are found in the
Indo-Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...
region. ::Feeding habits :::These species eat other
gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. T ...
s including cones. * Subgenus ''Endemoconus'' Iredale, 1931 ::Shell characters (living and fossil species) :::The shell is turbinate with a scalariform spire. The
protoconch A protoconch (meaning first or earliest or original shell) is an embryonic or larval shell which occurs in some classes of molluscs, e.g., the initial chamber of an ammonite or the larval shell of a gastropod. In older texts it is also called ...
is either paucispiral or multispiral. Whorl tops are concave and do not have cords. Nodules may be absent or may be present and persist on all whorls. A dentiform plait is present. The anal notch is shallow, and the anterior notch is absent. The
periostracum The periostracum ( ) is a thin, organic coating (or "skin") that is the outermost layer of the shell of many shelled animals, including molluscs and brachiopods. Among molluscs, it is primarily seen in snails and clams, i.e. in gastropods and ...
is tufted and the operculum is small to moderate in size. ::Radular tooth (not known for fossil species) :::The anterior section of the
radula The radula (, ; plural radulae or radulas) is an anatomical structure used by molluscs for feeding, sometimes compared to a tongue. It is a minutely toothed, chitinous ribbon, which is typically used for scraping or cutting food before the food ...
is much longer than the posterior section. The blade is short, a basal spur is absent, and the barb is short. The radular tooth has no serrations. An accessory process is present and is a short, simple triangular terminating cusp. ::Geographical distribution :::These species are found from Australia to New Zealand and in Madagascar. ::Feeding habits :::Unknown, however these species are believed to be
piscivorous A piscivore () is a carnivorous animal that eats primarily fish. The name ''piscivore'' is derived . Piscivore is equivalent to the Greek-derived word ichthyophage, both of which mean "fish eater". Fish were the diet of early tetrapod evoluti ...
based upon Bayesian cladistics analysis.


Species list

This list of species is based on the information in the
World Register of Marine Species The World Register of Marine Species (WoRMS) is a taxonomic database that aims to provide an authoritative and comprehensive list of names of marine organisms. Content The content of the registry is edited and maintained by scientific specialist ...
(
WoRMS Worms may refer to: *Worm, an invertebrate animal with a tube-like body and no limbs Places *Worms, Germany, a city **Worms (electoral district) *Worms, Nebraska, U.S. *Worms im Veltlintal, the German name for Bormio, Italy Arts and entertainme ...
) list. Species within the genus ''Endemoconus'' include: * ''Endemoconus bonfigliolii'' Bozzetti, 2010: synonym of ''
Conus bonfigliolii ''Conus bonfigliolii'' is a species of sea snail, a marine gastropod mollusc in the family Conidae, the cone snails and their allies.Bouchet, P. (2014). Conus bonfigliolii (Bozzetti, 2010). Accessed through: World Register of Marine Species ...
'' (Bozzetti, 2010) * ''Endemoconus boucheti'' (Richard, 1983): synonym of ''
Conasprella boucheti ''Conasprella boucheti'' is a species of sea snail, a marine gastropod mollusk in the family Conidae, the cone snails and their allies.Bouchet, P. (2015). Conasprella boucheti (Richard, 1983). Accessed through: World Register of Marine Speci ...
'' (Richard, 1983) * ''Endemoconus grohi'' (Tenorio & Poppe, 2004): synonym of ''
Conasprella grohi ''Conasprella grohi'' is a species of sea snail, a marine gastropod mollusk in the family Conidae, the cone snails and their allies.Bouchet, P. (2015). Conasprella grohi (Tenorio & Poppe, 2004). Accessed through: World Register of Marine Spe ...
'' (Tenorio & Poppe, 2004) * ''Endemoconus howelli'' (Iredale, 1929): synonym of '' Conasprella howelli'' Iredale, 1929 * ''Endemoconus ichinoseanus'' (Kuroda, 1956): synonym of ''
Conasprella ichinoseana ''Conasprella ichinoseana'' is a species of sea snail, a marine gastropod mollusk in the family Conidae, the cone snails and their allies.Bouchet, P. (2015). Conasprella ichinoseana (Kuroda, 1956). Accessed through: World Register of Marine Spec ...
'' (Kuroda, 1956) * ''Endemoconus ione'' (Fulton, 1938): synonym of ''
Conasprella ione ''Conasprella ione'', common name the ione cone, is a species of sea snail, a marine gastropod mollusk in the family Conidae, the cone snails and their allies.Bouchet, P. (2015). Conasprella ione (Fulton, 1938). Accessed through: World Register ...
'' (Fulton, 1938) * ''Endemoconus lozeti'' (Richard, 1980): synonym of ''
Conus lozeti ''Conus lozeti'' is a species of sea snail, a marine gastropod mollusk in the family Conidae, the cone snails and their allies. Like all species within the genus ''Conus'', these snails are predatory and venomous. They are capable of "stinging ...
'' Richard, 1980 * ''Endemoconus nadaensis'' Azuma & Toki, 1970: synonym of '' Conasprella articulata'' (G. B. Sowerby II, 1873) * ''Endemoconus otohimeae'' (Kuroda & Itô, 1961): synonym of ''
Conasprella otohimeae ''Conasprella otohimeae'', common name Otohime's cone, is a species of sea snail, a marine gastropod mollusk in the family Conidae, the cone snails and their allies. Like all species within the genus ''Conasprella'', these snails are predatory ...
'' (Kuroda & Itô, 1961) * ''Endemoconus raoulensis'' (Powell, 1958): synonym of '' Conasprella raoulensis'' Powell, 1958 * ''Endemoconus sculletti'' (Marsh, 1962): synonym of '' Conus sculletti'' Marsh, 1962 * ''Endemoconus sieboldii'' (Reeve, 1848): synonym of ''
Conasprella sieboldii ''Conasprella sieboldii'', common name Siebold's cone, is a species of sea snail, a marine gastropod mollusk in the family Conidae, the cone snails and their allies. Like all species within the genus ''Conasprella'', these cone snails are preda ...
'' (Reeve, 1848) * ''Endemoconus somalicus'' (Bozzetti, 2013): synonym of ''
Conasprella somalica ''Conasprella somalica'' is a species of sea snail, a marine gastropod mollusk in the family Conidae, the cone snails and their allies.Bouchet, P. (2015). Conasprella somalica (Bozzetti, 2013). Accessed through: World Register of Marine Species a ...
'' (Bozzetti, 2013) * ''Endemoconus spirofilis'' (Habe & Kosuge, 1970): synonym of ''
Conasprella spirofilis ''Conasprella spirofilis'' is a species of sea snail, a marine gastropod mollusk in the family Conidae, the cone snails and their allies. Like all species within the genus ''Conasprella'', these cone snails are predatory and venomous. They are ...
'' (Habe & Kosuge, 1970)


Significance of "alternative representation"

Prior to 2009, all species within the family
Conidae Conidae, with the current common name of "cone snails", is a taxonomic family (previously subfamily) of predatory sea snails, marine gastropod molluscs in the superfamily Conoidea. The 2014 classification of the superfamily Conoidea, groups onl ...
were placed in one genus, ''
Conus ''Conus'' is a genus of predatory sea snails, or cone snails, marine gastropod mollusks in the family Conidae.Bouchet, P.; Gofas, S. (2015). Conus Linnaeus, 1758. In: MolluscaBase (2015). Accessed through: World Register of Marine Species at ...
''. In 2009 however, J.K. Tucker and M.J. Tenorio proposed a classification system for the over 600 recognized
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
that were in the family. Their classification proposed 3 distinct families and 82 genera for the living species of
cone snails A cone is a three-dimensional geometric shape that tapers smoothly from a flat base (frequently, though not necessarily, circular) to a point called the apex or vertex. A cone is formed by a set of line segments, half-lines, or lines co ...
. This classification was based upon shell
morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines *Morphology (archaeology), study of the shapes or forms of artifacts *Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies, ...
,
radula The radula (, ; plural radulae or radulas) is an anatomical structure used by molluscs for feeding, sometimes compared to a tongue. It is a minutely toothed, chitinous ribbon, which is typically used for scraping or cutting food before the food ...
r differences,
anatomy Anatomy () is the branch of biology concerned with the study of the structure of organisms and their parts. Anatomy is a branch of natural science that deals with the structural organization of living things. It is an old science, having its ...
,
physiology Physiology (; ) is the scientific study of functions and mechanisms in a living system. As a sub-discipline of biology, physiology focuses on how organisms, organ systems, individual organs, cells, and biomolecules carry out the chemical ...
,
cladistics Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived char ...
, with comparisons to molecular (DNA) studies. Published accounts of genera within the
Conidae Conidae, with the current common name of "cone snails", is a taxonomic family (previously subfamily) of predatory sea snails, marine gastropod molluscs in the superfamily Conoidea. The 2014 classification of the superfamily Conoidea, groups onl ...
that include the genus ''Endemoconus'' include J.K. Tucker & M.J. Tenorio (2009), and Bouchet et al. (2011). Testing in order to try to understand the molecular phylogeny of the
Conidae Conidae, with the current common name of "cone snails", is a taxonomic family (previously subfamily) of predatory sea snails, marine gastropod molluscs in the superfamily Conoidea. The 2014 classification of the superfamily Conoidea, groups onl ...
was initially begun by Christopher Meyer and Alan Kohn, and is continuing, particularly with the advent of nuclear DNA testing in addition to mDNA testing. However, in 2011, some experts still prefer to use the traditional classification, where all species are placed in ''
Conus ''Conus'' is a genus of predatory sea snails, or cone snails, marine gastropod mollusks in the family Conidae.Bouchet, P.; Gofas, S. (2015). Conus Linnaeus, 1758. In: MolluscaBase (2015). Accessed through: World Register of Marine Species at ...
'' within the single family
Conidae Conidae, with the current common name of "cone snails", is a taxonomic family (previously subfamily) of predatory sea snails, marine gastropod molluscs in the superfamily Conoidea. The 2014 classification of the superfamily Conoidea, groups onl ...
: for example, according to the current November 2011 version of the
World Register of Marine Species The World Register of Marine Species (WoRMS) is a taxonomic database that aims to provide an authoritative and comprehensive list of names of marine organisms. Content The content of the registry is edited and maintained by scientific specialist ...
, all species within the family Conidae are in the genus ''
Conus ''Conus'' is a genus of predatory sea snails, or cone snails, marine gastropod mollusks in the family Conidae.Bouchet, P.; Gofas, S. (2015). Conus Linnaeus, 1758. In: MolluscaBase (2015). Accessed through: World Register of Marine Species at ...
''. The binomial names of species in the 82 cone snail genera listed in Tucker & Tenorio 2009 are recognized by the
World Register of Marine Species The World Register of Marine Species (WoRMS) is a taxonomic database that aims to provide an authoritative and comprehensive list of names of marine organisms. Content The content of the registry is edited and maintained by scientific specialist ...
as "alternative representations." Debate within the scientific community regarding this issue continues, and additional molecular phylogeny studies are being carried out in an attempt to clarify the issue.B. Vallejo, Jr. (2005), ''Inferring the mode of speciation in the Indo-West Pacific Conus (Gastropoda: Conidae)'', Journal of Biogeography 32:1429-1439. All this has been superseded in 2015 by the new classification of the Conidae


References


Further reading

* Kohn A. A. (1992). ''Chronological Taxonomy of ''Conus'', 1758-1840". Smithsonian Institution Press, Washington and London. * Monteiro A. (ed.) (2007)
The Cone Collector 1
1-28. * Berschauer D. (2010). ''Technology and the Fall of the Mono-Generic Family'
The Cone Collector 15
pp. 51-54 * Puillandre N., Meyer C.P., Bouchet P., and Olivera B.M. (2011), ''Genetic divergence and geographical variation in the deep-water Conus orbignyi complex (Mollusca: Conoidea)'', Zoologica Scripta 40(4) 350-363.


External links


To World Register of Marine Species

Gastropods.com: ''Conidae'' setting forth the genera recognized therein.

Gastropods.com: Image of ''Endemoconus bonfigliolii'' Bozzetti, L., 2010
{{Taxonbar, from=Q5376044 Conidae Gastropod subgenera