Encrinus Liliiformis
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''Encrinus'' is an
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
crinoid Crinoids are marine animals that make up the class Crinoidea. Crinoids that are attached to the sea bottom by a stalk in their adult form are commonly called sea lilies, while the unstalked forms are called feather stars or comatulids, which are ...
s, and "one of the most famous". It lived during the
Late Silurian The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the shortest period of the Paleozo ...
-
Late Triassic The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch ...
, and its
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
s have been found in
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
.


History

Fossils of ''Encrinus'' went by several names in
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
before the establishment of modern paleontology. In
Lower Saxony Lower Saxony (german: Niedersachsen ; nds, Neddersassen; stq, Läichsaksen) is a German state (') in northwestern Germany. It is the second-largest state by land area, with , and fourth-largest in population (8 million in 2021) among the 16 ...
, they were called ''Sonnenräder'' ("sun wheels"), while in
Thuringia Thuringia (; german: Thüringen ), officially the Free State of Thuringia ( ), is a state of central Germany, covering , the sixth smallest of the sixteen German states. It has a population of about 2.1 million. Erfurt is the capital and larg ...
and
Hesse Hesse (, , ) or Hessia (, ; german: Hessen ), officially the State of Hessen (german: links=no, Land Hessen), is a States of Germany, state in Germany. Its capital city is Wiesbaden, and the largest urban area is Frankfurt. Two other major histor ...
they were called ''Bonifatiuspfennige'' ("
Saint Boniface Boniface, OSB ( la, Bonifatius; 675 – 5 June 754) was an English Benedictines, Benedictine monk and leading figure in the Anglo-Saxon mission to the Germanic parts of the Frankish Empire during the eighth century. He organised significant ...
's pennies"). In southwestern Germany, they went by ''Hexengeld'' ("witches' money"). The animal was correctly reconstructed for the first time in 1729, although various parts of the animal had been described before that. ''Encrinus'' was described in 1764 by Johann Gerhard Reinhard Andreae. It was assigned to the order Encrinida by
Jack Sepkoski Joseph John Sepkoski Jr. (July 26, 1948 – May 1, 1999) was a University of Chicago paleontologist. Sepkoski studied the fossil record and the diversity of life on Earth. Sepkoski and David Raup contributed to the knowledge of extinction events. ...
in 2002.


Description

''Encrinus'' possessed a large cup- or crown-like structure at the top of its body, which has been described as resembling "an unopened
tulip Tulips (''Tulipa'') are a genus of spring-blooming perennial herbaceous bulbiferous geophytes (having bulbs as storage organs). The flowers are usually large, showy and brightly coloured, generally red, pink, yellow, or white (usually in warm ...
." This "cup" could be between and long. The structure displayed "five-fold
symmetry Symmetry (from grc, συμμετρία "agreement in dimensions, due proportion, arrangement") in everyday language refers to a sense of harmonious and beautiful proportion and balance. In mathematics, "symmetry" has a more precise definit ...
" and had a slightly
concave Concave or concavity may refer to: Science and technology * Concave lens * Concave mirror Mathematics * Concave function, the negative of a convex function * Concave polygon, a polygon which is not convex * Concave set * The concavity In ca ...
base. It bore ten arms (although in some species, such as ''E. carnallis'', it bore 20), which in turn bore many small branches (pinnules). When opened, the arms and pinnules formed a kind of "feeding fan", used to catch prey such as
plankton Plankton are the diverse collection of organisms found in Hydrosphere, water (or atmosphere, air) that are unable to propel themselves against a Ocean current, current (or wind). The individual organisms constituting plankton are called plankt ...
. "Microscopic interpinnular channels" suggest that ''Encrinus'' may have been an active
filter feeder Filter feeders are a sub-group of suspension feeding animals that feed by straining suspended matter and food particles from water, typically by passing the water over a specialized filtering structure. Some animals that use this method of feedin ...
, creating its own currents by beating its
cilia The cilium, plural cilia (), is a membrane-bound organelle found on most types of eukaryotic cell, and certain microorganisms known as ciliates. Cilia are absent in bacteria and archaea. The cilium has the shape of a slender threadlike projecti ...
. It is known that the arms could shut tightly in life (most ''Encrinus'' fossils are found in this position), but it is difficult to estimate the greatest angle at which they could open. It is possible that specimens recorded as being widest open demonstrate the normal angle of opening in life. A tegmen ("small leathery dome") covered the mouth and topped the cup. Any particles of food that the animal caught would be passed down the arms and below the tegmen via grooves, eventually leading to cilia, which in turn led to the mouth. When viewed from above, the stem was circular in shape. Large and small
ossicle The ossicles (also called auditory ossicles) are three bones in either middle ear that are among the smallest bones in the human body. They serve to transmit sounds from the air to the fluid-filled labyrinth (cochlea). The absence of the auditory ...
s covered the stem, creating a pattern of frequent small ossicles sandwiched between the large ones. The stem did not possess any
cirri Giovanni Battista Cirri (1 October 1724 – 11 June 1808) was an Italian cellist and composer in the 18th century. Biography Cirri was born in Forlì in the Emilia-Romagna Region of Italy. He had his first musical training with his brother ...
, and may have been held in place by roots of some sort. The breakage of the stem during the animal's life was a common occurrence, and it often survived long enough to form a new end on its shortened stem. Being from the Middle Triassic, ''Encrinus'' is the last known from its group of crinoids, as most others died out during the
Permian–Triassic extinction event The Permian–Triassic (P–T, P–Tr) extinction event, also known as the Latest Permian extinction event, the End-Permian Extinction and colloquially as the Great Dying, formed the boundary between the Permian and Triassic geologic periods, as ...
.


See also

*


References


External links

* {{Taxonbar, from=Q3725079 Prehistoric crinoid genera Triassic crinoids Middle Triassic animals of Europe