Empidonax Alnorum
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The alder flycatcher (''Empidonax alnorum'') is a small insect-eating
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
of the
tyrant flycatcher The tyrant flycatchers (Tyrannidae) are a family of passerine birds which occur throughout North and South America. They are considered the largest family of birds known to exist in the world, with more than 400 species. They are the most dive ...
family. The genus name ''
Empidonax The genus ''Empidonax'' is a group of small insect, insect-eating passerine birds in the tyrant flycatcher family, the Tyrannidae. The genus name ''Empidonax'' is from Ancient Greek ''empis'', "gnat", and ''anax'', "master". Most of these birds ...
'' is from
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic peri ...
''empis'', "
gnat A gnat () is any of many species of tiny flying insects in the dipterid suborder Nematocera, especially those in the families Mycetophilidae, Anisopodidae and Sciaridae. They can be both biting and non-biting. Most often they fly in large num ...
", and ''anax'', "master". The specific ''alnorum'' is
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
and means "of the
alder Alders are trees comprising the genus ''Alnus'' in the birch family Betulaceae. The genus comprises about 35 species of monoecious trees and shrubs, a few reaching a large size, distributed throughout the north temperate zone with a few sp ...
s".


Description

''E. alnorum'' is one of the larger ''Empidonax'' species, attaining a length of , and a mass of . The wingspan ranges from . Upperparts of ''E. alnorum'' are dull greenish-olive. The crown is a slightly darker olive-green than the back. The throat is white, with a darker breast band. Thin white rings surround the eyes. The bill is wide and moderate in length, with pinkish or yellow-orange lower mandible and black upper mandible. Wings are generally black, with white wing bars and white edges of the innermost secondaries (tertials). Juveniles have brownish upperparts, yellow underparts, and yellowish brown or buff wingbars.All about birds nternet(2015). Ithaca (NY): Cornell University; pdated 2015; cited 2016 Sept 13 Available from: https://www.allaboutbirds.org/guide/Alder_Flycatcher/lifehistory ''E. alnorum'' is similar in appearance to the eastern wood-peewee (''Contopus virens'') and the eastern phoebe (''Sayornis phoebe''), as well as the nearly identical willow flycatcher (''Empidonax traillii'') but ''E. alnorum'' can be distinguished by its smaller size and upward tail flicking behavior. Birds in the genus ''Empidonax'' are notoriously hard to identify. Novel approaches to identification have been employed to try and increase the accuracy of species identification of mist-netted individuals, such as using ratios of and differences between the measured lengths of various body parts, including wings and beaks. Vocalizations are often relied upon for identification to species in the field when birds cannot be manipulated by hand.Lowther PE. 1999. Alder Flycatcher (''Empidonax alnorum''). The Birds of North America (P. G. Rodewald, Ed.). Ithaca (NY): Cornell Lab of Ornithology; Retrieved from the Birds of North America: https://birdsna.org/Species-Account/bna/species/aldfly


Taxonomy

The alder flycatcher is sibling species with the willow flycatcher (''E. trailli''). The two species were grouped together as one species, the Traill's flycatcher, until 1973, when differences in vocalizations and habitat use lead to their acknowledgement as distinct species, which has since been supported by genetic data.Areta J, Mangini G, Gandoy F, Gorleri F, Gomez D, Depino E, Jordan E. 2016. Ecology and behavior of Alder Flycatchers (''Empidonax alnorum'') on their wintering grounds in Argentina. The Wilson journal of ornithology. 128(4): 830-845. A study analyzing the genetic makeup of 12 ''Empidonax'' species found that the Alder and Willow Flycatcher are each other's closest relatives. The Tyrant flycatcher genus is closely related to the Wood-Pewee genus (''Contopus''), and a hybrid of the two genera was reported to have been caught in Idaho.


Habitat and distribution

The summer breeding range of the species covers most of Canada and Alaska, descending to a southernmost point in the northeastern United States. In autumn, ''E. alnorum'' migrates south through the eastern United States, Mexico and Central America, wintering in the western range of South America. In the summer breeding season, ''E. alnorum'' occupies wet, dense, shrubby thickets of alder (''Alnus'' sp.), maple (''Acer'' sp.), and birch (''Betula'' sp.) at elevations below . During migration, the species occupies humid and semi-arid habitats, including forest edges and fields at elevations up to . Early successional scrubby vegetation or woodland edges are preferred habitats in winter, particularly those near water and below .Audubon guide to North American birds nternet N.d. National Audubon Society; ited 2017 Sept 13 Available from: http://www.audubon.org/field-guide/bird/alder-flycatcher


Behavior


Vocalizations

The vocalizations of the alder flycatcher are very useful for its identification in the field. The song is a distinct "fee-bee-o"; birds have been observed to throw their heads back and shake their tails while singing. There are a variety of calls generated by ''E. alnorum'', the most common of which is a "pit" sound produced when foraging. Other calls include those associated with aggressive or territorial behavior, including "double-peak", "zwee-oo", and "wee-oo" calls, and those linked with excitement, such as a "kitter" call.


Diet

''E. alnorum'' diet is composed primarily of insects such as those from the Hymenoptera (sawflies, bees, wasps and ants), Coleoptera (beetles), Diptera (flies), Lepidoptera (butterflies and moths) and Orthoptera (locusts, grasshoppers and crickets) families. Insects are captured during flight or gleaned from foliage of trees. In the winter, some birds will include fruit and seeds in their diet.


Reproduction

The alder flycatcher breeds in wet thickets of maple, alder and birch. Nests are positioned low in bushes within shrubby thickets, and are cup-shaped, built loosely of vegetative materials such as grass, weeds, pieces of bark, and small twigs. Strips of grass or bark can often be seen dangling from the bottom of the nest, and the inside is lined with soft materials such as plant down. Females primarily engage in nest building activities. Male ''E. alnorum'' vocalize to defend their breeding territory. As the species has not been extensively studied, courtship behavior is uncertain, but is believed to involve males chasing females through the trees. 3–4 eggs are laid per breeding season, and are creamy-white or buff in color and speckled with dark markings near the larger end of the egg. The female incubates the eggs for 12–14 days. Hatchlings are
altricial In biology, altricial species are those in which the young are underdeveloped at the time of birth, but with the aid of their parents mature after birth. Precocial species are those in which the young are relatively mature and mobile from the mome ...
and covered with tiny spots of olive-brown down. Both the male and female are involved in caring for the young. Juveniles take their first flight around 13–14 days of age.


Conservation status

''E. alnorum'' is a species of least concern according to the IUCN Redlist. Population numbers are stable in the United States, but have declined in Canada by approximately 44% in the period 1966–2014. The species has a rating of 9 out of 20 on the Continental Concern Score, and is a U.S.-Canada Stewardship species.


References


External links


Alder Flycatcher - ''Empidonax alnorum''
- USGS Patuxent Bird Identication InfoCenter

- Cornell Lab of Ornithology
Alder Flycatcher
Boreal Songbird Initiative {{Taxonbar, from=Q681933
alder flycatcher The alder flycatcher (''Empidonax alnorum'') is a small insect-eating bird of the tyrant flycatcher family. The genus name ''Empidonax'' is from Ancient Greek ''empis'', "gnat", and ''anax'', "master". The specific ''alnorum'' is Latin and means ...
Birds of Canada Birds of North America Birds of Appalachia (United States)
alder flycatcher The alder flycatcher (''Empidonax alnorum'') is a small insect-eating bird of the tyrant flycatcher family. The genus name ''Empidonax'' is from Ancient Greek ''empis'', "gnat", and ''anax'', "master". The specific ''alnorum'' is Latin and means ...
alder flycatcher The alder flycatcher (''Empidonax alnorum'') is a small insect-eating bird of the tyrant flycatcher family. The genus name ''Empidonax'' is from Ancient Greek ''empis'', "gnat", and ''anax'', "master". The specific ''alnorum'' is Latin and means ...