![Stinger of wild Pandinus imperator](https://upload.wikimedia.org/wikipedia/commons/7/78/Stinger_of_wild_Pandinus_imperator.jpg)
The emperor scorpion, ''Pandinus imperator'', is a
species
In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of
scorpion
Scorpions are predatory arachnids of the order Scorpiones. They have eight legs, and are easily recognized by a pair of grasping pincers and a narrow, segmented tail, often carried in a characteristic forward curve over the back and always end ...
native to rainforests and savannas in
West Africa
West Africa or Western Africa is the westernmost region of Africa. The United Nations defines Western Africa as the 16 countries of Benin, Burkina Faso, Cape Verde, The Gambia, Ghana, Guinea, Guinea-Bissau, Ivory Coast, Liberia, Mali, Maurit ...
. It is one of the largest scorpions in the world and lives for 6–8 years. Its body is black, but like other scorpions it glows pastel green or blue under
ultraviolet light
Ultraviolet (UV) is a form of electromagnetic radiation with wavelength from 10 nm (with a corresponding frequency around 30 PHz) to 400 nm (750 THz), shorter than that of visible light, but longer than X-rays. UV radiation i ...
. It is a popular species in the
pet trade
Wildlife trade refers to the of products that are derived from non-domesticated animals or plants usually extracted from their natural environment or raised under controlled conditions. It can involve the trade of living or dead individuals, ti ...
, and is protected by
CITES
CITES (shorter name for the Convention on International Trade in Endangered Species of Wild Fauna and Flora, also known as the Washington Convention) is a multilateral treaty to protect endangered plants and animals from the threats of interna ...
to prevent over-collecting that might impact the species' survival.
Description
![Emperor scorpion or Imperial scorpion (Pandinus imperator)](https://upload.wikimedia.org/wikipedia/commons/6/6d/Emperor_scorpion_or_Imperial_scorpion_%28Pandinus_imperator%29.jpg)
The emperor scorpion (''Pandinus imperator'') is one of the largest species of scorpion in the world, with adults averaging about in length and a weight of 30 g. However, some species of forest scorpions are fairly similar to the emperor scorpion in size, and one scorpion, ''
Heterometrus swammerdami
''Gigantometrus swammerdami'', commonly called the giant forest scorpion, is a scorpion belonging to the family Scorpionidae. It is native to India. It is the world's largest scorpion species with 23 cm (9 in) in length, and weigh as m ...
'', holds the record for being the world's largest scorpion at 9 inches (23 cm) in length. The large pincers are blackish-red and have a granular texture. The front part of the body, or
prosoma
The cephalothorax, also called prosoma in some groups, is a tagma of various arthropods, comprising the head and the thorax fused together, as distinct from the abdomen behind. (The terms ''prosoma'' and ''opisthosoma'' are equivalent to ''cepha ...
, is made up of four sections, each with a pair of legs. Behind the fourth pair of legs are comb-like structures known as
pectines, which tend to be longer in males than in females. The tail, known as the
metasoma
The metasoma is the posterior part of the body, or tagma, of arthropods whose body is composed of three parts, the other two being the prosoma and the mesosoma. In insects, it contains most of the digestive tract, respiratory system, and circula ...
, is long and curves back over the body. It ends in the large receptacle containing the venom glands and is tipped with a sharp, curved
stinger
A stinger (or sting) is a sharp organ found in various animals (typically insects and other arthropods) capable of injecting venom, usually by piercing the epidermis of another animal.
An insect sting is complicated by its introduction of v ...
. Scorpion stings can be categorized as mild (similar to a bee sting) to severe to humans depending on the species. Most people are not severely affected by the emperor scorpion's sting, though some people may be allergic to scorpion stings in general. Sensory hairs cover the pincers and tail, enabling the emperor scorpion to detect prey through vibrations in the air and ground.
When gravid (pregnant), the body of a female expands to expose the whitish membranes connecting the segments. The emperor scorpion fluoresces greenish-blue under
ultra-violet
Ultraviolet (UV) is a form of electromagnetic radiation with wavelength from 10 nm (with a corresponding frequency around 30 PHz) to 400 nm (750 THz), shorter than that of visible light, but longer than X-rays. UV radiation i ...
light.
[ Accessed October 20, 2011.]
They are known for their docile behavior and almost harmless sting; they do not use their sting to defend themselves when they are adults, however, they may use it in their adolescent stages. They prefer to use their pincers to crush and dismember their prey. Their exoskeleton is very sclerotic, causing them to have a metallic greenish-black color.
Emperor scorpions are often confused with a similar genus (''Heterometrus''), and are one of the most famous scorpions.
Different
ion channel toxins
An ion () is an atom or molecule with a net electrical charge.
The charge of an electron is considered to be negative by convention and this charge is equal and opposite to the charge of a proton, which is considered to be positive by convent ...
have been isolated from the venom of the emperor scorpion, including Pi1, Pi2,
Pi3,
Pi4 and Pi7.
Habitat and distribution
The emperor scorpion is an African rainforest species, but also present in
savanna
A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground to ...
. It is found in a number of African countries, including
Benin
Benin ( , ; french: Bénin , ff, Benen), officially the Republic of Benin (french: République du Bénin), and formerly Dahomey, is a country in West Africa. It is bordered by Togo to the west, Nigeria to the east, Burkina Faso to the north ...
,
Burkina Faso
Burkina Faso (, ; , ff, 𞤄𞤵𞤪𞤳𞤭𞤲𞤢 𞤊𞤢𞤧𞤮, italic=no) is a landlocked country in West Africa with an area of , bordered by Mali to the northwest, Niger to the northeast, Benin to the southeast, Togo and Ghana to the ...
,
Côte d’Ivoire
Ivory Coast, also known as Côte d'Ivoire, officially the Republic of Côte d'Ivoire, is a country on the southern coast of West Africa. Its capital is Yamoussoukro, in the centre of the country, while its largest city and economic centre is ...
,
Gambia
The Gambia,, ff, Gammbi, ar, غامبيا officially the Republic of The Gambia, is a country in West Africa. It is the smallest country within mainland AfricaHoare, Ben. (2002) ''The Kingfisher A-Z Encyclopedia'', Kingfisher Publicatio ...
,
Ghana
Ghana (; tw, Gaana, ee, Gana), officially the Republic of Ghana, is a country in West Africa. It abuts the Gulf of Guinea and the Atlantic Ocean to the south, sharing borders with Ivory Coast in the west, Burkina Faso in the north, and To ...
,
Guinea
Guinea ( ),, fuf, 𞤘𞤭𞤲𞤫, italic=no, Gine, wo, Gine, nqo, ߖߌ߬ߣߍ߫, bm, Gine officially the Republic of Guinea (french: République de Guinée), is a coastal country in West Africa. It borders the Atlantic Ocean to the we ...
,
Guinea-Bissau
Guinea-Bissau ( ; pt, Guiné-Bissau; ff, italic=no, 𞤘𞤭𞤲𞤫 𞤄𞤭𞤧𞤢𞥄𞤱𞤮, Gine-Bisaawo, script=Adlm; Mandinka: ''Gine-Bisawo''), officially the Republic of Guinea-Bissau ( pt, República da Guiné-Bissau, links=no ) ...
,
Togo
Togo (), officially the Togolese Republic (french: République togolaise), is a country in West Africa. It is bordered by Ghana to the west, Benin to the east and Burkina Faso to the north. It extends south to the Gulf of Guinea, where its c ...
,
Liberia
Liberia (), officially the Republic of Liberia, is a country on the West African coast. It is bordered by Sierra Leone to Liberia–Sierra Leone border, its northwest, Guinea to its north, Ivory Coast to its east, and the Atlantic Ocean ...
,
Mali
Mali (; ), officially the Republic of Mali,, , ff, 𞤈𞤫𞤲𞥆𞤣𞤢𞥄𞤲𞤣𞤭 𞤃𞤢𞥄𞤤𞤭, Renndaandi Maali, italics=no, ar, جمهورية مالي, Jumhūriyyāt Mālī is a landlocked country in West Africa. Mali ...
,
Nigeria
Nigeria ( ), , ig, Naìjíríyà, yo, Nàìjíríà, pcm, Naijá , ff, Naajeeriya, kcg, Naijeriya officially the Federal Republic of Nigeria, is a country in West Africa. It is situated between the Sahel to the north and the Gulf o ...
,
Senegal
Senegal,; Wolof: ''Senegaal''; Pulaar: 𞤅𞤫𞤲𞤫𞤺𞤢𞥄𞤤𞤭 (Senegaali); Arabic: السنغال ''As-Sinighal'') officially the Republic of Senegal,; Wolof: ''Réewum Senegaal''; Pulaar : 𞤈𞤫𞤲𞤣𞤢𞥄𞤲𞤣𞤭 ...
,
Sierra Leone
Sierra Leone,)]. officially the Republic of Sierra Leone, is a country on the southwest coast of West Africa. It is bordered by Liberia to the southeast and Guinea surrounds the northern half of the nation. Covering a total area of , Sierra ...
and
Cameroon
Cameroon (; french: Cameroun, ff, Kamerun), officially the Republic of Cameroon (french: République du Cameroun, links=no), is a country in west-central Africa. It is bordered by Nigeria to the west and north; Chad to the northeast; the C ...
.
This species inhabits both
tropical forest
Tropical forests (a.k.a. jungle) are forested landscapes in tropical regions: ''i.e.'' land areas approximately bounded by the tropic of Cancer and Capricorn, but possibly affected by other factors such as prevailing winds.
Some tropical fores ...
and open savannas. The emperor scorpion burrows beneath the soil and hides beneath rocks and debris,
and also often burrows in
termite mound
Mound-building termites are a group of termite species that live in mounds. These termites live in Africa, Australia and South America. The mounds sometimes have a diameter of . Most of the mounds are in well-drained areas. Termite mounds usually ...
s.
Feeding habits
In the wild, emperor scorpions primarily consume insects and other terrestrial invertebrates, although termites constitute a large portion of their diet. Larger vertebrates, such as rodents and lizards, are occasionally eaten. Emperor scorpions will burrow through termite mounds up to 6 feet deep in order to hunt prey. Their large claws help in tearing apart prey while their tail stinger injects venom at the same time for liquifying food. Juveniles rely on their venomous sting to paralyze prey while adults use their large claws to tear apart prey.
Conservation and human impact
File:Emperor scorpion feeding.jpg, Young female emperor scorpion kept as a pet
File:Emperor Scorpion in hand John.jpg, ''P. imperator'' is very docile and tolerant of human contact.
African emperor scorpion
venom
Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
contains the toxins
imperatoxin and
pandinotoxin
Pandinotoxins are toxins from the venom of the emperor scorpion '' Pandinus imperator''. They are selective blockers of voltage-gated potassium channels
Sources
The source for the pandinotoxins is the venom of the scorpion '' Pandinus imperat ...
.
Due to its docile nature, large size, and hardiness, ''P. imperator'' is a popular
scorpion
Scorpions are predatory arachnids of the order Scorpiones. They have eight legs, and are easily recognized by a pair of grasping pincers and a narrow, segmented tail, often carried in a characteristic forward curve over the back and always end ...
in the pet trade, which has led to such over-collecting in the wild that it is now a
CITES
CITES (shorter name for the Convention on International Trade in Endangered Species of Wild Fauna and Flora, also known as the Washington Convention) is a multilateral treaty to protect endangered plants and animals from the threats of interna ...
-listed
animal
Animals are multicellular, eukaryotic organisms in the Kingdom (biology), biological kingdom Animalia. With few exceptions, animals Heterotroph, consume organic material, Cellular respiration#Aerobic respiration, breathe oxygen, are Motilit ...
.
They feed readily on crickets and worms available to keepers, and they can live up to 8 years in captivity.
Comparison with the Asian forest scorpion
![Baby Asian Forest scorpion](https://upload.wikimedia.org/wikipedia/commons/8/8e/Baby_Asian_Forest_scorpion.jpg)
Emperor scorpions are often confused with
Asian forest scorpions (''
Heterometrus
''Heterometrus'', whose members are also known by the collective vernacular name giant forest scorpions, is a genus of scorpions belonging to the family Scorpionidae. It is distributed widely across tropical and subtropical southeastern Asia, in ...
'' sp.) as the two species are indeed very similar in appearance. Both are very popular as pets due to their impressive size, mild venom, and usually docile nature. However, and while this may vary, Asian forest scorpions tend to be more high-strung and defensive than emperor scorpions, and noticeable differences exist between the two in the claws and stinger. Emperors have rough pincers, which also tend to be shorter but thicker than those of their Asian counterparts. Asian forest scorpions lack the red coloration that emperor scorpion pincers often exhibit, and their telson and aculeus are black, while emperor scorpions tend to have yellow to reddish yellow telson as juveniles that turn red as they mature. Even babies exhibit a distinct visual difference, as emperor scorpion scorplings usually show quite visible red pigmentation in their pincers, which is completely absent in baby Asian forest scorpions.
References
External links
*
*
{{DEFAULTSORT:Emperor Scorpion
Scorpionidae
Scorpions of Africa
Animals described in 1842
Taxa named by Carl Ludwig Koch