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(Marie) Louis Emberger (23 January 1897 – 30 November 1969) was a French
botanist Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek wo ...
and phytogeographer, at the
University of Montpellier The University of Montpellier (french: Université de Montpellier) is a public university, public research university located in Montpellier, in south-east of France. Established in 1220, the University of Montpellier is one of the oldest univ ...
.


Life

Emberger was born at Thann, in
Haut-Rhin Haut-Rhin (, ; Alsatian: ''Owerelsàss'' or '; german: Oberelsass, ) is a department in the Grand Est region of France, bordering both Germany and Switzerland. It is named after the river Rhine. Its name means ''Upper Rhine''. Haut-Rhin is the ...
, France in 1897, which was then part of German occupied Alsace. He developed an interest in Natural History, exploring the
Rhine plain The Upper Rhine Plain, Rhine Rift Valley or Upper Rhine Graben (German: ''Oberrheinische Tiefebene'', ''Oberrheinisches Tiefland'' or ''Oberrheingraben'', French: ''Vallée du Rhin'') is a major rift, about and on average , between Basel in the s ...
of Alsace, and the nearby
Vosges mountains The Vosges ( , ; german: Vogesen ; Franconian and gsw, Vogese) are a range of low mountains in Eastern France, near its border with Germany. Together with the Palatine Forest to the north on the German side of the border, they form a single ...
. At the age of 17, to avoid conscription into the German army, he escaped to
Lyons Lyon,, ; Occitan: ''Lion'', hist. ''Lionés'' also spelled in English as Lyons, is the third-largest city and second-largest metropolitan area of France. It is located at the confluence of the rivers Rhône and Saône, to the northwest of th ...
, in France. There he began studies in biologie at the
University of Lyons The University of Lyon (french: Université de Lyon), located in Lyon and Saint-Étienne, France, is a center for higher education and research comprising 11 members and 24 associated institutions. The three main universities in this center are: C ...
, and obtained a degree in ''sciences naturelles'' in 1918. He obtained his doctorate under Professor Marie Antoine Alexandre Guilliermond, then head of the Department of Agricultural Botany, at Lyons. Following his studies, his first position was as an
organic chemistry Organic chemistry is a subdiscipline within chemistry involving the scientific study of the structure, properties, and reactions of organic compounds and organic materials, i.e., matter in its various forms that contain carbon atoms.Clayden, J.; ...
technician in the Faculty of Medicine and Pharmacy. There he produced his first publication in 1919 in the '' Bulletin de la Société Botanique de France'', an account of his exploration at
Grande Chartreuse Grande Chartreuse () is the head monastery of the Carthusian religious order. It is located in the Chartreuse Mountains, north of the city of Grenoble, in the commune of Saint-Pierre-de-Chartreuse (Isère), France. History Originally, the ch ...
. He began working as a pharmacist in 1920 and within a year had had six papers presented at the ''
Académie des Sciences The French Academy of Sciences (French: ''Académie des sciences'') is a learned society, founded in 1666 by Louis XIV at the suggestion of Jean-Baptiste Colbert, to encourage and protect the spirit of French scientific research. It was at the ...
'' by
Gaston Bonnier Gaston Eugène Marie Bonnier (9 April 1853 in Paris – 2 January 1922) was a French botanist and plant ecologist. Biography Bonnier first studied at École Normale Supérieure in Paris from 1873 to 1876. Together with Charles Flahault, he st ...
. In 1921 he was appointed as a lecturer in the Faculty of Pharmacy at the
University of Montpellier The University of Montpellier (french: Université de Montpellier) is a public university, public research university located in Montpellier, in south-east of France. Established in 1220, the University of Montpellier is one of the oldest univ ...
. Within five years he had been appointed head of the botany department at the ''Institut scientifique de Rabat'' in
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to ...
(1926–1936). Returning to France he was, for a short time, professor at the Faculty of Science at
Clermont-Ferrand Clermont-Ferrand (, ; ; oc, label=Auvergnat (dialect), Auvergnat, Clarmont-Ferrand or Clharmou ; la, Augustonemetum) is a city and Communes of France, commune of France, in the Auvergne-Rhône-Alpes regions of France, region, with a population ...
, before once more taking a position in Montpellier, succeeding his step-father,
Charles Flahault Charles Henri Marie Flahault (3 October 1852 – 3 February 1935) was a French botanist, among the early pioneers of phytogeography, phytosociology, and forest ecology. The word '' relevé'' for a plant community sample is his invention. Early li ...
as head of the botany department (Directeur de l'Institut botanique de l'Université et du Centre d'Etudes phytosocio-logiques de Montpellier). There he worked closely with
Josias Braun-Blanquet Josias Braun-Blanquet (3 August 1884 – 20 September 1980) was an influential phytosociologist and botanist. Braun-Blanquet was born in Chur, Switzerland and died in Montpellier, France. Biography Work In Josias Braun-Blanquet's dissertation, s ...
, also a student of Flahault. He remained in that position until his death at St Sulpice in
Savoie Savoie (; Arpitan: ''Savouè'' or ''Savouè-d'Avâl''; English: ''Savoy'' ) is a department in the Auvergne-Rhône-Alpes region, Southeastern France. Located in the French Alps, its prefecture is Chambéry. In 2019, Savoie had a population of ...
in 1969.


Work

Emberger's scientific contributions cover four main areas of research,
cytology Cell biology (also cellular biology or cytology) is a branch of biology that studies the structure, function, and behavior of cells. All living organisms are made of cells. A cell is the basic unit of life that is responsible for the living and ...
,
biogeography Biogeography is the study of the distribution of species and ecosystems in geographic space and through geological time. Organisms and biological communities often vary in a regular fashion along geographic gradients of latitude, elevation, ...
,
comparative morphology Comparative anatomy is the study of similarities and differences in the anatomy of different species. It is closely related to evolutionary biology and phylogeny (the evolution of species). The science began in the classical era, continuing in t ...
and
phylogeny A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spec ...
, and
biosystematics Biological systematics is the study of the diversification of living forms, both past and present, and the Correlation and dependence, relationships among living things through time. Relationships are visualized as evolutionary trees (synonyms: c ...
. His cytological work focussed on the
ferns A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except th ...
,
horsetails ''Equisetum'' (; horsetail, snake grass, puzzlegrass) is the only living genus in Equisetaceae, a family of ferns, which reproduce by spores rather than seeds. ''Equisetum'' is a "living fossil", the only living genus of the entire subclass Eq ...
, and the
lycopods Lycopodiopsida is a class of vascular plants known as lycopods, lycophytes or other terms including the component lyco-. Members of the class are also called clubmosses, firmosses, spikemosses and quillworts. They have dichotomously branching s ...
. His biogeographical work concentrated on the vegetation of the
Mediterranean basin In biogeography, the Mediterranean Basin (; also known as the Mediterranean Region or sometimes Mediterranea) is the region of lands around the Mediterranean Sea that have mostly a Mediterranean climate, with mild to cool, rainy winters and w ...
and in particular the western
High Atlas High Atlas, also called the Grand Atlas ( ar, الأطلس الكبير, Al-Aṭlas al-Kabīr; french: Haut Atlas; shi, ⴰⴷⵔⴰⵔ ⵏ ⴷⵔⵏ ''Adrar n Dern''), is a mountain range in central Morocco, North Africa, the highest part of t ...
region of Morocco. He published work on the distribution and classification of Moroccan
flora Flora is all the plant life present in a particular region or time, generally the naturally occurring (indigenous) native plants. Sometimes bacteria and fungi are also referred to as flora, as in the terms '' gut flora'' or '' skin flora''. E ...
, in particular, halfah grass (''
Stipa tenacissima ''Stipa tenacissima'' (esparto, esparto grass, halfah grass, alfa grass, or needle grass) is a perennial grass of northwestern Africa and the southern part of the Iberian Peninsula. Distribution ''Stipa tenacissima'' is an endemic species of th ...
''). His Moroccan work also included studies on the
altitudinal zonation Altitudinal zonation (or elevational zonation) in mountainous regions describes the natural layering of ecosystems that occurs at distinct elevations due to varying environmental conditions. Temperature, humidity, soil composition, and solar radi ...
of the mountain regions there and the phytogeography of the
desert climate The desert climate or arid climate (in the Köppen climate classification ''BWh'' and ''BWk''), is a dry climate sub-type in which there is a severe excess of evaporation over precipitation. The typically bald, rocky, or sandy surfaces in desert ...
, in particular the argan tree (''
Argania spinosa ''Argania'' (Tashelhit: ⴰⵔⴳⴰⵏ ''Argan'') is a genus of flowering plants containing the sole species ''Argania spinosa'', known as argan, a tree endemic to the calcareous semidesert Sous valley of southwestern Morocco and to the regi ...
''). Other work included studies of
association Association may refer to: *Club (organization), an association of two or more people united by a common interest or goal *Trade association, an organization founded and funded by businesses that operate in a specific industry *Voluntary associatio ...
in equatorial forests, which led to a biogeographical classification of
climate Climate is the long-term weather pattern in an area, typically averaged over 30 years. More rigorously, it is the mean and variability of meteorological variables over a time spanning from months to millions of years. Some of the meteorologic ...
, in which he drew comparisons between Australia and homologous zones of the Mediterranean. In biosystematics he developed a
classification Classification is a process related to categorization, the process in which ideas and objects are recognized, differentiated and understood. Classification is the grouping of related facts into classes. It may also refer to: Business, organizat ...
scheme for
vascular plants Vascular plants (), also called tracheophytes () or collectively Tracheophyta (), form a large group of land plants ( accepted known species) that have lignified tissues (the xylem) for conducting water and minerals throughout the plant. They al ...
.


Selected publications

* * * * see also ''Taxon. Lit., ed. 2 (TL2)'',


Legacy

Louis Embeerger is the authority for 168
taxa In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...
, such as '' Agropyron pseudofestucoides'' Emb.


References


Bibliography

* * * * * * * * * * * {{DEFAULTSORT:Emberger, Louis 20th-century French botanists 1897 births 1969 deaths University of Montpellier faculty