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''Elornis'' is an extinct genus of
flamingo Flamingos or flamingoes are a type of Wader, wading bird in the Family (biology), family Phoenicopteridae, which is the only extant family in the order Phoenicopteriformes. There are four flamingo species distributed throughout the Americas ...
from the
Late Oligocene The Chattian is, in the geologic timescale, the younger of two ages or upper of two stages of the Oligocene Epoch/Series. It spans the time between . The Chattian is preceded by the Rupelian and is followed by the Aquitanian (the lowest stage ...
of Ronzon,
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
. Although the name was coined in the 1850s, the name remained a nomen nudum until later publications by French zoologist
Henri Milne-Edwards Henri Milne-Edwards (23 October 1800 – 29 July 1885) was an eminent French zoologist. Biography Henri Milne-Edwards was the 27th child of William Edwards, an English planter and colonel of the militia in Jamaica and Elisabeth Vaux, a Frenchw ...
. ''Elornis'' has been historically considered to be a member of the
Phoenicopteridae Flamingos or flamingoes are a type of wading bird in the family Phoenicopteridae, which is the only extant family in the order Phoenicopteriformes. There are four flamingo species distributed throughout the Americas (including the Caribbean) ...
, but more recent research suggests it may have been more basal. Research on this taxon is however difficult as the current whereabouts of the fossil material are unknown, limiting data to the description and illustrations of Milne-Edwards.


History

The name ''Elornis'' was first coined by
Auguste Aymard Auguste Aymard (5 December 1808 – 26 June 1889) was a French prehistorian and palaeontologist who lived and died in Puy-en-Velay (Haute-Loire). He described the fossil '' Entelodon magnus'' and the fossil genera ''Anancus'' and ''Amphechinus''. ...
in a report on the paleontology of the French
Haute Loire Haute-Loire (; oc, Naut Léger or ''Naut Leir''; English: Upper Loire) is a landlocked department in the Auvergne-Rhône-Alpes region of south-central France. Named after the Loire River, it is surrounded by the departments of Loire, Ardèche, ...
region, which was presented on two occasions to the Congres Seientifique de France in 1855. In his report, Aymard introduces several names for fossil birds, including ''Elornis grandis'', ''Elornis (?) littoralis'' and ''Elornis (?) antiquus''. However, very little information is given in the report, only stating that these birds are flamingos and neither giving a full description nor illustrating the material. This effectively renders the names as coined by Aymard absolute
nomina nuda In taxonomy, a ''nomen nudum'' ('naked name'; plural ''nomina nuda'') is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be one, but it has not been published with an adequate descr ...
. However, in spite of the names being coined under these circumstances, they found widespread use in subsequent years. Most notable among them was
Henri Milne-Edwards Henri Milne-Edwards (23 October 1800 – 29 July 1885) was an eminent French zoologist. Biography Henri Milne-Edwards was the 27th child of William Edwards, an English planter and colonel of the militia in Jamaica and Elisabeth Vaux, a Frenchw ...
, who discussed ''Elornis'' while also providing illustrations of the fossil material. Subsequent authors then proceeded to base their own works in regards to ''Elornis'' on Milne-Edwards' work, oftentimes without ever reading the paper in which the name was first coined, partially due to the rare nature of the publication. The
nomenclature Nomenclature (, ) is a system of names or terms, or the rules for forming these terms in a particular field of arts or sciences. The principles of naming vary from the relatively informal naming conventions, conventions of everyday speech to the i ...
of ''Elornis'' was eventually covered in greater detail by
Storrs Olson Storrs Lovejoy Olson (April 3, 1944 – January 20, 2021) was an American biologist and ornithologist who spent his career at the Smithsonian Institution, retiring in 2008. One of the world's foremost avian paleontologists, he was best known ...
, who wrote that due to Aymard's names being nomina nuda, the status of his taxa was entirely dependent on Milne-Edwards description. Thus, instead of Aymard being credited as the authority behind ''Elornis'', it would have to be Milne-Edwards. Two more problems arose during Olson's research on ''Elornis'' however. For one, Milne-Edwards' work was not published in a single publication, but rather over the course of several livraisons leaving a two year gap between the publishing of the ''Elornis'' illustrations and the eventual description via text. Furthermore, Aymard's original material was presumed lost, with the descriptions and illustrations given by Milne-Edwards being the only available data. ''Elornis littoralis'', as described by Milne-Edwards, was based on various postcranial remains including the
pelvis The pelvis (plural pelves or pelvises) is the lower part of the trunk, between the abdomen and the thighs (sometimes also called pelvic region), together with its embedded skeleton (sometimes also called bony pelvis, or pelvic skeleton). The ...
, tarsometatarsi,
tibiotarsi The tibiotarsus is the large bone between the femur and the tarsometatarsus in the leg of a bird. It is the fusion of the proximal part of the tarsus with the tibia. A similar structure also occurred in the Mesozoic Heterodontosauridae. These sma ...
and furculae. ''Elornis antiquus'' was intended to be based on a
humerus The humerus (; ) is a long bone in the arm that runs from the shoulder to the elbow. It connects the scapula and the two bones of the lower arm, the radius and ulna, and consists of three sections. The humeral upper extremity consists of a roun ...
by Aymard, however this idea was immediately discarded by Milne-Edwards who considered it to be synonymous with ''E. littoralis''. This would make the humerus part of the
syntype In biological nomenclature, a syntype is any one of two or more biological types that is listed in a description of a taxon where no holotype was designated. Precise definitions of this and related terms for types have been established as part of ...
material and not, as incorrectly suggested by Lydekker and Brodkorb, a
lectotype In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to anchor or centralizes the ...
. The material of ''Elornis grandis'' received less attention, the humerus Aymard intended as the holotype only mentioned to be "as big as that of a flamingo" by Milne-Edwards and not illustrated at all. Due to the broken up nature of the publication, the illustrations of ''Elornis littoralis'' were published before the text description of both ''E. littoralis'' and ''E. grandis''. The caption makes note of the name and in the absence of images of ''E. grandis'' on the same plate, ''Elornis littoralis'' would have to be the type species by
monotypy In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
, published several years prior to ''Elornis grandis''. One more species was named by Lydekker in 1891, designated ''Elornis (?) anglicus''. Later research however placed this species in the genus '' Actiornis'' and recovered it as an
ibis The ibises () (collective plural ibis; classical plurals ibides and ibes) are a group of long-legged wading birds in the family Threskiornithidae, that inhabit wetlands, forests and plains. "Ibis" derives from the Latin and Ancient Greek word f ...
.


Description

The humerus of ''Elornis littoralis'' was described as being approximately a quarter smaller than the humerus of a flamingo, while ''Elornis grandis'' was described as being of similar size. It is however not specified which species of flamingo is meant.


Classification

''Elornis'' was first recognized as a flamingo by Auguste Aymard when the name was first coined. Milne-Edwards later agreed with the assignment and Olson, although incapable of studying the fossil material itself, concurred with the identification on the basis of the illustrations. Subsequent papers repeat this identification, placing ''Elornis'' in the
Phoenicopteridae Flamingos or flamingoes are a type of wading bird in the family Phoenicopteridae, which is the only extant family in the order Phoenicopteriformes. There are four flamingo species distributed throughout the Americas (including the Caribbean) ...
. This is questioned by Mayr, who argues that ''Elornis'' would be better placed outside of ''Phoenicopteridae'' given the less derived morphology of other Oligocene flamingo relatives.


References

{{Taxonbar, from=Q16113575 Oligocene Phoenicopteridae Prehistoric bird genera Fossil taxa described in 1868 Fossils of France