Elm Sphinx
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''Ceratomia amyntor'', the elm sphinx or four-horned sphinx, is a North American
moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...
in the family Sphingidae. The species was first described by
Carl Geyer Peter Carl Friedrich Geyer (1802–1889) was a German entomologist who wrote and illustrated various supplements to Jacob Hübner's works on Lepidoptera. Carl Geyer was by profession an artist. He is not to be confused with Karl Andreas Geyer (180 ...
in 1835. It has a wingspan of - inches (8.2 - 11.5 cm). As the name suggests, the
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The ...
e (
caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder Sym ...
s) feed on elm trees (''
Ulmus Elms are deciduous and semi-deciduous trees comprising the flowering plant genus ''Ulmus'' in the plant family Ulmaceae. They are distributed over most of the Northern Hemisphere, inhabiting the temperate and tropical-montane regions of Nor ...
''), but they can also be found feeding on birch ('' Betula''), basswood (''
Tilia ''Tilia'' is a genus of about 30 species of trees or bushes, native throughout most of the temperateness, temperate Northern Hemisphere. The tree is known as linden for the European species, and basswood for North American species. In Britain a ...
''), and cherry (''
Prunus ''Prunus'' is a genus of trees and shrubs, which includes (among many others) the fruits plums, cherries, peaches, nectarines, apricots, and almonds. Native to the North American temperate regions, the neotropics of South America, and the p ...
''). When the caterpillars are ready, they crawl to the bottom of the
host A host is a person responsible for guests at an event or for providing hospitality during it. Host may also refer to: Places * Host, Pennsylvania, a village in Berks County People *Jim Host (born 1937), American businessman * Michel Host ...
tree, where they crawl underneath the soil and pupate and may overwinter underground if late enough into the year. Vegetable growers should be aware of this larvae due to its insatiable appetite. One of these larvae are capable in devouring huge amounts of plant's foliage and even succulent stems.


Range

''C. amyntor'' can be found from
Nova Scotia Nova Scotia ( ; ; ) is one of the thirteen provinces and territories of Canada. It is one of the three Maritime provinces and one of the four Atlantic provinces. Nova Scotia is Latin for "New Scotland". Most of the population are native Eng ...
west to
Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...
and western
North Dakota North Dakota () is a U.S. state in the Upper Midwest, named after the Native Americans in the United States, indigenous Dakota people, Dakota Sioux. North Dakota is bordered by the Canadian provinces of Saskatchewan and Manitoba to the north a ...
and
Colorado Colorado (, other variants) is a state in the Mountain West subregion of the Western United States. It encompasses most of the Southern Rocky Mountains, as well as the northeastern portion of the Colorado Plateau and the western edge of t ...
; south to central
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and to ...
, the Gulf Coast,
Texas Texas (, ; Spanish language, Spanish: ''Texas'', ''Tejas'') is a state in the South Central United States, South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2 ...
,
Louisiana Louisiana , group=pronunciation (French: ''La Louisiane'') is a state in the Deep South and South Central regions of the United States. It is the 20th-smallest by area and the 25th most populous of the 50 U.S. states. Louisiana is borde ...
and
New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe , LargestCity = Albuquerque , LargestMetro = Tiguex , OfficialLang = None , Languages = English, Spanish ( New Mexican), Navajo, Ker ...
.


Status

The species is not threatened.


Life cycle

''C amyntor'' adults fly as a single brood in the northern portions of their range from June to July. There are two broods further south, flying later into the year, and five broods have been confirmed in Louisiana from March to October.


Egg

Tiny, translucent, lime-green eggs deposited on the underside of host leaves. The shells are transparent and pearly after hatching.


Larva

Larvae come in two different color phases; a green phase, and a dark phase which differs in shades of brown, orange, and somewhat of a pinkish brown. Larvae complete all five
instar An instar (, from the Latin '' īnstar'', "form", "likeness") is a developmental stage of arthropods, such as insects, between each moult (''ecdysis''), until sexual maturity is reached. Arthropods must shed the exoskeleton in order to grow or ass ...
s within approximately one month. The first instar is the same shade of green as its egg. As the larva progresses through its instars, change in structure is noticeable. The caterpillar sports four, long, horn-like projections - in addition to the tail horn - from above its head, which is why the elm sphinx is sometimes called the four-horned sphinx.


Pupa

As with other Sphingidae, ''C. amyntor'' goes through a "wandering" phase where it stops feeding and burrows into the soil in order to pupate. Before pupating, the larva shrinks a considerable amount and then sheds its remaining skin that distinguishes it as a caterpillar, revealing its shell-like pupa.


Imago

The overall coloring of the
imago In biology, the imago (Latin for "image") is the last stage an insect attains during its metamorphosis, its process of growth and development; it is also called the imaginal stage, the stage in which the insect attains maturity. It follows the f ...
, or adult, elm sphinx is a gray-brown with wavy markings covering the
forewing Insect wings are adult outgrowths of the insect exoskeleton that enable insects to fly. They are found on the second and third thoracic segments (the mesothorax and metathorax), and the two pairs are often referred to as the forewings and hindwin ...
s. In the upper center of the forewings, as with other '' Ceratomia'' species, there is a small white dot. The
hindwing Insect wings are adult outgrowths of the insect exoskeleton that enable insects to fly. They are found on the second and third thoracic segments (the mesothorax and metathorax), and the two pairs are often referred to as the forewings and hindwi ...
s are a plain gray with a darker band running the length of the near-edge. The outer ends of the forewings and hindwings are both dashed with a light-dark-light pattern. Three broken, dark bands run the length of the body from the thorax to the tip of the abdomen. The antennae are stiff and wiry and have hook at the tips. Some specimens have a prominent splash of bright white on their upper head and wings and stand out considerably from other ''C. amyntor''. Ceratomia amyntor MHNT CUT 2010 0 406 Chaffeys Locks Leeds Co Ontario, male dorsal.jpg, ''Ceratomia amyntor'' ♂ Ceratomia amyntor MHNT CUT 2010 0 406 Chaffeys Locks Leeds Co Ontario, male ventral.jpg, ''Ceratomia amyntor'' ♂ △ Ceratomia amyntor MHNT CUT 2010 0 406 Delaware, Ontario, female dorsal.jpg, ''Ceratomia amyntor'' ♀ Ceratomia amyntor MHNT CUT 2010 0 406 Delaware, Ontario, female ventral.jpg, ''Ceratomia amyntor'' ♀ △


Food plants

*''Ulmus'' (
elm Elms are deciduous and semi-deciduous trees comprising the flowering plant genus ''Ulmus'' in the plant family Ulmaceae. They are distributed over most of the Northern Hemisphere, inhabiting the temperate and tropical-montane regions of North ...
) *''Betula'' (
birch A birch is a thin-leaved deciduous hardwood tree of the genus ''Betula'' (), in the family Betulaceae, which also includes alders, hazels, and hornbeams. It is closely related to the beech-oak family Fagaceae. The genus ''Betula'' contains 30 ...
) *''Tilia'' (
basswood ''Tilia americana'' is a species of tree in the family Malvaceae, native to eastern North America, from southeast Manitoba east to New Brunswick, southwest to northeast Oklahoma, southeast to South Carolina, and west along the Niobrara River to ...
) *''Prunus'' (
cherry A cherry is the fruit of many plants of the genus ''Prunus'', and is a fleshy drupe (stone fruit). Commercial cherries are obtained from cultivars of several species, such as the sweet ''Prunus avium'' and the sour ''Prunus cerasus''. The nam ...
) *
Tomato The tomato is the edible berry of the plant ''Solanum lycopersicum'', commonly known as the tomato plant. The species originated in western South America, Mexico, and Central America. The Mexican Nahuatl word gave rise to the Spanish word ...
plants *
Pepper Pepper or peppers may refer to: Food and spice * Piperaceae or the pepper family, a large family of flowering plant ** Black pepper * ''Capsicum'' or pepper, a genus of flowering plants in the nightshade family Solanaceae ** Bell pepper ** Chili ...
plants


See also

*
Lepidoptera Lepidoptera ( ) is an order (biology), order of insects that includes butterfly, butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 Family (biology), families and 46 Taxonomic r ...
*
Moths Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...
* Sphingidae


References

* Villiard, P. (1969). ''Moths and How to Rear Them''. New York: Funk & Wagnalls.
LCCN The Library of Congress Control Number (LCCN) is a serially based system of numbering cataloged records in the Library of Congress, in the United States. It is not related to the contents of any book, and should not be confused with Library of ...
68-27516


External links

*
"''Ceratomia amyntor'' The Elm Sphinx or Four-horned Sphinx (Geyer, [1835])"
''Sphingidae of the Americas''. Archived October 7, 2007. {{DEFAULTSORT:Ceratomia Amyntor Ceratomia Moths described in 1835 Moths of North America