Elephant cognition is
animal cognition
Animal cognition encompasses the mental capacities of non-human animals including insect cognition. The study of animal conditioning and learning used in this field was developed from comparative psychology. It has also been strongly influenc ...
as present in
elephants
Elephants are the largest existing land animals. Three living species are currently recognised: the African bush elephant, the African forest elephant, and the Asian elephant. They are the only surviving members of the family Elephantidae and ...
. Most contemporary
ethologists
Ethology is the scientific study of animal behaviour, usually with a focus on behaviour under natural conditions, and viewing behaviour as an evolutionarily adaptive trait. Behaviourism as a term also describes the scientific and objectiv ...
view the elephant as one of the world's most intelligent animals. With a mass of just over , an elephant's brain has more mass than that of any other land animal, and although the largest
whale
Whales are a widely distributed and diverse group of fully aquatic placental marine mammals. As an informal and colloquial grouping, they correspond to large members of the infraorder Cetacea, i.e. all cetaceans apart from dolphins and ...
s have body masses twenty times those of a typical elephant, a whale's brain is barely twice the mass of an elephant's brain. In addition, elephants have around 257 billion
neurons
A neuron, neurone, or nerve cell is an electrically excitable cell that communicates with other cells via specialized connections called synapses. The neuron is the main component of nervous tissue in all animals except sponges and placozoa. N ...
.
[ Elephant brains are similar to humans' and many other mammals' in terms of general connectivity and functional areas, with several unique structural differences. Although initially estimated to have as many neurons as a human brain,] the elephant's cortex
Cortex or cortical may refer to:
Biology
* Cortex (anatomy), the outermost layer of an organ
** Cerebral cortex, the outer layer of the vertebrate cerebrum, part of which is the ''forebrain''
*** Motor cortex, the regions of the cerebral cortex i ...
has about one-third of the number of neurons as a human brain.
Elephants manifest a wide variety of behaviors, including those associated with grief
Grief is the response to loss, particularly to the loss of someone or some living thing that has died, to which a bond or affection was formed. Although conventionally focused on the emotional response to loss, grief also has physical, cogni ...
, learning, mimicry
In evolutionary biology, mimicry is an evolved resemblance between an organism and another object, often an organism of another species. Mimicry may evolve between different species, or between individuals of the same species. Often, mimicry f ...
, play, altruism
Altruism is the principle and moral practice of concern for the welfare and/or happiness of other human beings or animals, resulting in a quality of life both material and spiritual. It is a traditional virtue in many cultures and a core as ...
, use of tools, compassion
Compassion motivates people to go out of their way to relieve the physical, mental or emotional pains of others and themselves. Compassion is often regarded as being sensitive to the emotional aspects of the suffering of others. When based on n ...
, cooperation
Cooperation (written as co-operation in British English) is the process of groups of organisms working or acting together for common, mutual, or some underlying benefit, as opposed to working in competition for selfish benefit. Many animal a ...
, self-awareness
In philosophy of self, self-awareness is the experience of one's own personality or individuality. It is not to be confused with consciousness in the sense of qualia. While consciousness is being aware of one's environment and body and lifesty ...
, memory
Memory is the faculty of the mind by which data or information is encoded, stored, and retrieved when needed. It is the retention of information over time for the purpose of influencing future action. If past events could not be remembered, ...
, and communication
Communication (from la, communicare, meaning "to share" or "to be in relation with") is usually defined as the transmission of information. The term may also refer to the message communicated through such transmissions or the field of inquir ...
. Further evidence suggests elephants may understand pointing
Pointing is a gesture specifying a direction from a person's body, usually indicating a location, person, event, thing or idea. It typically is formed by extending the arm, hand, and index finger, although it may be functionally similar to other ...
: the ability to nonverbally communicate an object by extending a finger, or equivalent. It is thought they are equal with cetaceans and primate
Primates are a diverse order of mammals. They are divided into the strepsirrhines, which include the lemurs, galagos, and lorisids, and the haplorhines, which include the tarsiers and the simians (monkeys and apes, the latter including huma ...
s in this regard. Due to such claims of high intelligence and due to strong family ties of elephants, some researchers argue it is morally wrong for humans to cull
In biology, culling is the process of segregating organisms from a group according to desired or undesired characteristics. In animal breeding, it is the process of removing or segregating animals from a breeding stock based on a specific tr ...
them. Aristotle
Aristotle (; grc-gre, Ἀριστοτέλης ''Aristotélēs'', ; 384–322 BC) was a Greek philosopher and polymath during the Classical period in Ancient Greece. Taught by Plato, he was the founder of the Peripatetic school of phil ...
described the elephant as "the animal that surpasses all others in wit and mind
The mind is the set of faculties responsible for all mental phenomena. Often the term is also identified with the phenomena themselves. These faculties include thought, imagination, memory, will, and sensation. They are responsible for various m ...
."
Structure of the brain
Cerebral cortex
The elephant (both Asian
Asian may refer to:
* Items from or related to the continent of Asia:
** Asian people, people in or descending from Asia
** Asian culture, the culture of the people from Asia
** Asian cuisine, food based on the style of food of the people from Asi ...
and African
African or Africans may refer to:
* Anything from or pertaining to the continent of Africa:
** People who are native to Africa, descendants of natives of Africa, or individuals who trace their ancestry to indigenous inhabitants of Africa
*** Ethn ...
) has a very large and highly complex neocortex
The neocortex, also called the neopallium, isocortex, or the six-layered cortex, is a set of layers of the mammalian cerebral cortex involved in higher-order brain functions such as sensory perception, cognition, generation of motor commands, sp ...
, a trait also shared by humans
Humans (''Homo sapiens'') are the most abundant and widespread species of primate, characterized by bipedalism and exceptional cognitive skills due to a large and complex brain. This has enabled the development of advanced tools, culture, ...
, apes
Apes (collectively Hominoidea ) are a clade of Old World simians native to sub-Saharan Africa and Southeast Asia (though they were more widespread in Africa, most of Asia, and as well as Europe in prehistory), which together with its sister g ...
and certain dolphin
A dolphin is an aquatic mammal within the infraorder Cetacea. Dolphin species belong to the families Delphinidae (the oceanic dolphins), Platanistidae (the Indian river dolphins), Iniidae (the New World river dolphins), Pontoporiidae (the ...
species.
Asian elephants have the greatest volume of cerebral cortex
The cerebral cortex, also known as the cerebral mantle, is the outer layer of neural tissue of the cerebrum of the brain in humans and other mammals. The cerebral cortex mostly consists of the six-layered neocortex, with just 10% consisting of ...
available for cognitive
Cognition refers to "the mental action or process of acquiring knowledge and understanding through thought, experience, and the senses". It encompasses all aspects of intellectual functions and processes such as: perception, attention, thought, ...
processing of all existing land animals. It exceeds that of any primate
Primates are a diverse order of mammals. They are divided into the strepsirrhines, which include the lemurs, galagos, and lorisids, and the haplorhines, which include the tarsiers and the simians (monkeys and apes, the latter including huma ...
species, with one study suggesting elephants be placed in the category of great apes
The Hominidae (), whose members are known as the great apes or hominids (), are a taxonomic family of primates that includes eight extant species in four genera: '' Pongo'' (the Bornean, Sumatran and Tapanuli orangutan); ''Gorilla'' (the east ...
in terms of cognitive abilities for tool use and tool making.
The elephant brain exhibits a gyral pattern more complex and with more numerous convolutions, or brain folds, than that of humans, other primates
Primates are a diverse order of mammals. They are divided into the strepsirrhines, which include the lemurs, galagos, and lorisids, and the haplorhines, which include the tarsiers and the simians (monkeys and apes, the latter including huma ...
, or carnivores
A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose food and energy requirements derive from animal tissues (mainly muscle, fat and other sof ...
, but less complex than that of cetaceans. Elephants are believed to rank equal with dolphins in terms of problem-solving abilities, and many scientists tend to rank elephant intelligence
Intelligence has been defined in many ways: the capacity for abstraction, logic, understanding, self-awareness, learning, emotional knowledge, reasoning, planning, creativity, critical thinking, and problem-solving. More generally, it can b ...
at the same level as cetaceans; a 2011 article published by ABC Science
ABC News, or ABC News and Current Affairs, is a public news service produced by the Australian Broadcasting Corporation. Broadcasting within Australia and the rest of the world, the service covers both local and world affairs.
The division of ...
suggests that, "elephants re assmart as chimps, nddolphins
A dolphin is an aquatic mammal within the infraorder Cetacea. Dolphin species belong to the families Delphinidae (the oceanic dolphins), Platanistidae (the Indian river dolphins), Iniidae (the New World river dolphins), Pontoporiidae (t ...
".[
]
Other areas of the brain
Elephants also have a very large and highly convoluted hippocampus
The hippocampus (via Latin from Greek , 'seahorse') is a major component of the brain of humans and other vertebrates. Humans and other mammals have two hippocampi, one in each side of the brain. The hippocampus is part of the limbic system, a ...
, a brain structure in the limbic system
The limbic system, also known as the paleomammalian cortex, is a set of brain structures located on both sides of the thalamus, immediately beneath the medial temporal lobe of the cerebrum primarily in the forebrain.Schacter, Daniel L. 2012. ''Ps ...
that is much bigger than that of any human, primate
Primates are a diverse order of mammals. They are divided into the strepsirrhines, which include the lemurs, galagos, and lorisids, and the haplorhines, which include the tarsiers and the simians (monkeys and apes, the latter including huma ...
or cetacean
Cetacea (; , ) is an infraorder of aquatic mammals that includes whales, dolphins, and porpoises. Key characteristics are their fully aquatic lifestyle, streamlined body shape, often large size and exclusively carnivorous diet. They propel them ...
. The hippocampus of an elephant takes up about 0.7% of the central structures of the brain, comparable to 0.5% for humans and with 0.1% in Risso's dolphin
Risso's dolphin (''Grampus griseus'') is a dolphin, the only species of the genus ''Grampus''. Some of the closest related species to these dolphins include: pilot whales (''Globicephala'' spp.), pygmy killer whales (''Feresa attenuata''), melon ...
s and 0.05% in bottlenose dolphins.
The hippocampus is linked to emotion
Emotions are mental states brought on by neurophysiological changes, variously associated with thoughts, feelings, behavioral responses, and a degree of pleasure or displeasure. There is currently no scientific consensus on a definition. ...
through the processing of certain types of memory
Memory is the faculty of the mind by which data or information is encoded, stored, and retrieved when needed. It is the retention of information over time for the purpose of influencing future action. If past events could not be remembered, ...
, especially spatial. This is thought to be possibly why elephants suffer from psychological flashbacks and the equivalent of post-traumatic stress disorder
Post-traumatic stress disorder (PTSD) is a mental and behavioral disorder that can develop because of exposure to a traumatic event, such as sexual assault, warfare, traffic collisions, child abuse, domestic violence, or other threats on ...
(PTSD).
The encephalization quotient
Encephalization quotient (EQ), encephalization level (EL), or just encephalization is a relative brain size measure that is defined as the ratio between observed to predicted brain mass for an animal of a given size, based on nonlinear regressi ...
(EQ) (the size of the brain relative to body size) of elephants ranges from 1.13 to 2.36. The average EQ is 2.14 for Asian elephants, and 1.67 for African, with the overall average being 1.88. In comparison to other animals, the La Plata
La Plata () is the capital city of Buenos Aires Province, Argentina. According to the , it has a population of 654,324 and its metropolitan area, the Greater La Plata, has 787,294 inhabitants. It is located 9 kilometers (6 miles) inland from th ...
dolphin has an EQ of 1.67; the Ganges river dolphin
The Ganges river dolphin (''Platanista gangetica'') is a species of toothed whale classified in the family Platanistidae. It lives in the Ganges and related rivers of South Asia, namely in the countries of India, Nepal, and Bangladesh. It is rel ...
of 1.55; the orca
The orca or killer whale (''Orcinus orca'') is a toothed whale belonging to the oceanic dolphin family, of which it is the largest member. It is the only Extant taxon, extant species in the genus ''Orcinus'' and is recognizable by its black ...
of 2.57; the bottlenose dolphin
Bottlenose dolphins are aquatic mammals in the genus ''Tursiops.'' They are common, cosmopolitan members of the family Delphinidae, the family of oceanic dolphins. Molecular studies show the genus definitively contains two species: the common ...
of 4.14; and the tucuxi dolphin
The tucuxi (''Sotalia fluviatilis''), alternatively known in Peru ''bufeo gris'' or ''bufeo negro'', is a species of freshwater dolphin found in the rivers of the Amazon basin. The word ''tucuxi'' is derived from the Tupi language word ''tuchu ...
of 4.56; chimpanzees at 2.49; dogs at 1.17, cats at 1.00; and mice at 0.50. Humans have an EQ of 7.44.
Brain size at birth relative to adult brain size
Comparing brain size at birth to the size of a fully developed adult's brain is one way to estimate how much an animal relies on learning as opposed to instinct. The majority of mammals are born with a brain close to 90% of the adult weight, while humans are born with 28%,[ ]bottlenose dolphins
Bottlenose dolphins are aquatic mammals in the genus ''Tursiops.'' They are common, cosmopolitan members of the family Delphinidae, the family of oceanic dolphins. Molecular studies show the genus definitively contains two species: the comm ...
with 42.5%, chimpanzee
The chimpanzee (''Pan troglodytes''), also known as simply the chimp, is a species of great ape native to the forest and savannah of tropical Africa. It has four confirmed subspecies and a fifth proposed subspecies. When its close relative th ...
s with 54%,[ and elephants with 35%.] This may indicate that elephants require the second highest amount of learning while developing (next to humans), and that their behavior is less instinctual than taught. This is further supported by the elephant's long juvenile period and large temporal lobes
The temporal lobe is one of the four major lobes of the cerebral cortex in the brain of mammals. The temporal lobe is located beneath the lateral fissure on both cerebral hemispheres of the mammalian brain.
The temporal lobe is involved in pro ...
, which are associated in the storage of memories.
Spindle neurons
Spindle cells
Von Economo neurons (VENs), also called spindle neurons, are a specific class of mammalian cortical neurons characterized by a large spindle-shaped soma (or body) gradually tapering into a single apical axon (the ramification that ''transmits ...
appear to play a central role in the development of intelligent behavior. As well as in humans and the rest of the great apes, spindle neurons are also found in the brains of both Asian and African elephants, as well as humpback whale
The humpback whale (''Megaptera novaeangliae'') is a species of baleen whale. It is a rorqual (a member of the family Balaenopteridae) and is the only species in the genus ''Megaptera''. Adults range in length from and weigh up to . The hu ...
s, fin whale
The fin whale (''Balaenoptera physalus''), also known as finback whale or common rorqual and formerly known as herring whale or razorback whale, is a cetacean belonging to the parvorder of baleen whales. It is the second-longest species of cet ...
s, killer whale
The orca or killer whale (''Orcinus orca'') is a toothed whale belonging to the oceanic dolphin family, of which it is the largest member. It is the only extant species in the genus ''Orcinus'' and is recognizable by its black-and-white pa ...
s, sperm whale
The sperm whale or cachalot (''Physeter macrocephalus'') is the largest of the toothed whales and the largest toothed predator. It is the only living member of the genus ''Physeter'' and one of three extant species in the sperm whale famil ...
s, bottlenose dolphins, Risso's dolphin
Risso's dolphin (''Grampus griseus'') is a dolphin, the only species of the genus ''Grampus''. Some of the closest related species to these dolphins include: pilot whales (''Globicephala'' spp.), pygmy killer whales (''Feresa attenuata''), melon ...
s, and beluga whale
The beluga whale () (''Delphinapterus leucas'') is an Arctic and sub-Arctic cetacean. It is one of two members of the family Monodontidae, along with the narwhal, and the only member of the genus ''Delphinapterus''. It is also known as the whi ...
s. The remarkable similarity between the elephant brain and the human brain supports the thesis of convergent evolution
Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last com ...
.
Elephant society
The elephant has one of the most closely knit societies of any living species. Elephant families can only be separated by death or capture. Cynthia Moss
Cynthia Jane Moss (born July 24, 1940) is an American ethologist and conservationist, wildlife researcher, and writer. Her studies have concentrated on the demography, behavior, social organization, and population dynamics of the African elephant ...
, an ethologist
Ethology is the scientific study of animal behaviour, usually with a focus on behaviour under natural conditions, and viewing behaviour as an evolutionarily adaptive trait. Behaviourism as a term also describes the scientific and objectiv ...
specialising in elephants, recalls an event involving a family of African elephants:
::Two members of the family were shot by poachers, who were subsequently chased off by the remaining elephants. Although one of the elephants died, the other, named Tina, remained standing, but with knees beginning to give way. Two family members, Trista and Teresia (Tina's mother), walked to both sides of Tina and leaned in to hold her up. Eventually, Tina grew so weak, she fell to the ground and died. However, Trista and Teresia did not give up but continually tried to lift her. They managed to get Tina into a sitting position, but her body was lifeless and fell to the ground again. As the other elephant family members became more intensely involved in the aid, they tried to put grass into Tina's mouth. Teresia then put her tusks beneath Tina's head and front quarters and proceeded to lift her. As she did so, her right tusk broke completely off, right up to the lip and nerve cavity. The elephants gave up trying to lift Tina but did not leave her; instead, they began to bury her in a shallow grave and throw leaves over her body. They stood over Tina for the night and then began to leave in the morning. The last to leave was Teresia.
Because elephants are so closely knit and highly matriarchal, a family can be devastated by the death of another (especially a matriarch), and some groups never recover their organization. Cynthia Moss has observed a mother, after the death of her calf, walk sluggishly at the back of a family for many days.[
]Edward Topsell
Edward Topsell (''circa'' 1572 – 1625) was an English cleric and author best remembered for his bestiary.
Topsell was born and educated in Sevenoaks, Kent. He attended Christ's College, Cambridge, earned his B.A. and probably an M.A., as well, ...
stated in his publication ''The History of Four-Footed Beasts'' in 1607, "There is no creature among all the Beasts of the world which hath so great and ample demonstration of the power and wisdom of almighty God as the elephant." Elephants are believed to be on par with chimpanzees with regard to their cooperative skills.
Elephant altruism
Elephants are thought to be highly altruistic
Altruism is the principle and moral practice of concern for the welfare and/or happiness of other human beings or animals, resulting in a quality of life both material and spiritual. It is a traditional virtue in many cultures and a core asp ...
animals that even aid other species, including humans, in distress. In India, an elephant was helping locals lift logs by following a truck and placing the logs in pre-dug holes upon instruction from the mahout
A mahout is an elephant rider, trainer, or keeper. Mahouts were used since antiquity for both civilian and military use. Traditionally, mahouts came from ethnic groups with generations of elephant keeping experience, with a mahout retaining h ...
(elephant trainer). At a certain hole, the elephant refused to lower the log. The mahout came to investigate the hold-up and noticed a dog sleeping in the hole. The elephant only lowered the log when the dog was gone. When an elephant is hurt, other elephants (even if they are unrelated) aid them.[
Moss has often seen elephants going out of their way to avoid hurting or killing a human, even when it was difficult for them (such as having to walk backwards to avoid a person). Joyce Poole documented an encounter told to her by Colin Francombe on Kuki Gallman's Laikipia Ranch. A ranch herder was walking alongside camels when he came across a family of elephants. The ]matriarch
Matriarchy is a social system in which women hold the primary power positions in roles of authority. In a broader sense it can also extend to moral authority, social privilege and control of property.
While those definitions apply in general E ...
charged at him and knocked him over with her trunk, breaking one of his legs. In the evening, when he did not return, a search party was sent in a truck to find him. When the party discovered him, he was being guarded by an elephant. The animal charged the truck, so they shot over her and scared her away. The herdsman later told them that when he could not stand up, the elephant used her trunk to lift him under the shade of a tree. She guarded him for the day and would gently touch him with her trunk.[
]
Self-medication
Elephants in Africa self-medicate by chewing on the leaves of a tree from the family Boraginaceae
Boraginaceae, the borage or forget-me-not family, includes about 2,000 species of shrubs, trees and herbs in 146, to 156 genera with a worldwide distribution.
The APG IV system from 2016 classifies the Boraginaceae as single family of the or ...
, which induces labour
Labour or labor may refer to:
* Childbirth, the delivery of a baby
* Labour (human activity), or work
** Manual labour, physical work
** Wage labour, a socioeconomic relationship between a worker and an employer
** Organized labour and the labour ...
. Kenyans also use this tree for the same purpose.
Death ritual
Scientists often debate the extent to which elephants feel emotion.[ Elephants have been one of few species of mammals other than '']Homo sapiens sapiens
Human taxonomy is the classification of the human species (systematic name ''Homo sapiens'', Latin: "wise man") within zoological taxonomy. The systematic genus, ''Homo'', is designed to include both anatomically modern humans and extinct varie ...
'' and Neanderthals
Neanderthals (, also ''Homo neanderthalensis'' and erroneously ''Homo sapiens neanderthalensis''), also written as Neandertals, are an Extinction, extinct species or subspecies of archaic humans who lived in Eurasia until about 40,000 years ag ...
known to have or have had any recognizable ritual around death. Elephants show a keen interest in the bones of their own kind (even unrelated elephants that have died long ago). They are often seen gently investigating the bones with their trunks and feet while remaining very quiet. Sometimes elephants that are completely unrelated to the deceased still visit their graves.[
Elephant researcher ]Martin Meredith
Martin Meredith is a historian, journalist, and biographer. He has written several books on Africa and its modern history.
Meredith first worked as a foreign correspondent in Africa for ''The Observer'' and ''Sunday Times'', then as a research ...
recalls in his book an occurrence of a typical elephant death ritual as witnessed by Anthony Hall-Martin, a South African biologist who had studied elephants in Addo, South Africa, for over eight years. The entire family of a dead matriarch
Matriarchy is a social system in which women hold the primary power positions in roles of authority. In a broader sense it can also extend to moral authority, social privilege and control of property.
While those definitions apply in general E ...
, including her young calf, were all gently touching her body with their trunks, trying to lift her. The elephant herd were all rumbling loudly. The calf was observed to be weeping and made sounds that sounded like a scream, but then the entire herd fell silent. They then began to throw leaves and dirt over the body and broke off tree branches to cover her. They spent the next two days quietly standing over her body. They sometimes left to get water or food, but they would always return.
Occurrences of elephants' behaving this way around human beings are common throughout Africa. On many occasions, they have buried dead or sleeping humans or aided them when they were hurt.[
Meredith also recalls an event told to him by ]George Adamson
George Alexander Graham Adamson MBE (3 February 1906 – 20 August 1989), also known as the ''Baba ya Simba'' ("Father of Lions" in Swahili), was a Kenyan wildlife conservationist and author. He and his wife, Joy, were depicted in the film ''B ...
, a Kenyan game warden, regarding an old Turkana woman who fell asleep under a tree after losing her way home. When she woke up, there was an elephant standing over her, gently touching her. She kept very still because she was very frightened. As other elephants arrived, they began to scream loudly and buried her under branches. She was found the next morning by the local herdsmen, unharmed.[
George Adamson also recalls when he shot a ]bull
A bull is an intact (i.e., not castrated) adult male of the species ''Bos taurus'' (cattle). More muscular and aggressive than the females of the same species (i.e., cows), bulls have long been an important symbol in many religions,
includin ...
elephant from a herd that kept breaking into the government gardens of northern Kenya. George gave the elephant's meat to local Turkana tribesmen and then dragged the rest of the carcass half a mile (800 m) away. That night, the other elephants found the body and took the shoulder blade and leg bone and returned the bones to the exact spot the elephant was killed.
Play
Poole has observed wild African elephants at play on many occasions. They apparently do things for their own and others' entertainment. Elephants have been seen sucking up water, holding their trunk high in the air, and then spraying the water like a fountain.[
]
Mimicry
Recent studies have shown that elephants can also mimic
MIMIC, known in capitalized form only, is a former simulation computer language developed 1964 by H. E. Petersen, F. J. Sansom and L. M. Warshawsky of Systems Engineering Group within the Air Force Materiel Command at the Wright-Patterson AFB in ...
sounds they hear. The discovery was found when Mlaika, an orphaned elephant, would copy the sound of trucks passing by. So far, the only other animal
Animals are multicellular, eukaryotic organisms in the Kingdom (biology), biological kingdom Animalia. With few exceptions, animals Heterotroph, consume organic material, Cellular respiration#Aerobic respiration, breathe oxygen, are Motilit ...
s that are thought to mimic
MIMIC, known in capitalized form only, is a former simulation computer language developed 1964 by H. E. Petersen, F. J. Sansom and L. M. Warshawsky of Systems Engineering Group within the Air Force Materiel Command at the Wright-Patterson AFB in ...
sounds are whales
Whales are a widely distributed and diverse group of fully aquatic placental marine mammals. As an informal and colloquial grouping, they correspond to large members of the infraorder Cetacea, i.e. all cetaceans apart from dolphins and ...
, dolphins
A dolphin is an aquatic mammal within the infraorder Cetacea. Dolphin species belong to the families Delphinidae (the oceanic dolphins), Platanistidae (the Indian river dolphins), Iniidae (the New World river dolphins), Pontoporiidae (t ...
, bat
Bats are mammals of the order Chiroptera.''cheir'', "hand" and πτερόν''pteron'', "wing". With their forelimbs adapted as wings, they are the only mammals capable of true and sustained flight. Bats are more agile in flight than most ...
s, primates
Primates are a diverse order of mammals. They are divided into the strepsirrhines, which include the lemurs, galagos, and lorisids, and the haplorhines, which include the tarsiers and the simians (monkeys and apes, the latter including huma ...
and birds
Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
.
Calimero, an African elephant who was 23 years old, also exhibited a unique form of mimicry. He was in a Swiss zoo with some Asian elephants. Asian elephants use chirps that are different from African elephants' deep rumbling noises. Calimero also began to chirp and not make the deep calls that his species normally would.
Kosik, an Indian elephant
The Indian elephant (''Elephas maximus indicus'') is one of four extant recognised subspecies of the Asian elephant and native to mainland Asia.
Since 1986, the Asian elephant has been listed as Endangered on the IUCN Red List as the wild po ...
at Everland
Everland () is South Korea's largest theme park. Located at the Everland Resort in Yongin, a city in Gyeonggi-do, it receives 5.85 million visitors annually and was ranked nineteenth in the world for amusement park attendance in 2018. As of 2 ...
Amusement Park, South Korea
South Korea, officially the Republic of Korea (ROK), is a country in East Asia, constituting the southern part of the Korea, Korean Peninsula and sharing a Korean Demilitarized Zone, land border with North Korea. Its western border is formed ...
can imitate up to five Korean words, including ''sit'', ''no'', ''yes'' and ''lie down''. Kosik produces these human-like sounds by putting his trunk in his mouth and then shaking it while breathing out, similar to how people whistle with their fingers.
Ecologist O’Connell-Rodwell’s conducted research in 1997 which concluded that elephants create low-frequency vibrations (seismic signals) through their trunks and feet to communicate across long distances. Elephants use contact calls to stay in touch with one another when they are out of one another's sight. In 2004, Joseph Soltis conducted a study to understand the low-frequency vocalization elephants used to communicate across short-distances. The research found that closely allied female elephants were more likely to produce 'rumbles' to other members at twice the rate of those who had lesser integrated members. Female elephants are able to remember and distinguish the contact calls of female family and bond group members from those of females outside of their extended family network. They can also distinguish between the calls of family units depending upon how frequently they came across them.
Poole, who is part of the Amboseli Elephant Research Project
The Amboseli Elephant Research Project is a long-term research project on the ethology of the African elephant, operated by the nonprofit Amboseli Trust for Elephants. The project studies the elephant's social behavior, age structure and populati ...
in Kenya, has demonstrated vocal learning and imitation in elephants of sounds made by each other and in the environment. She is beginning to research whether sounds made by elephants have dialect
The term dialect (from Latin , , from the Ancient Greek word , 'discourse', from , 'through' and , 'I speak') can refer to either of two distinctly different types of Linguistics, linguistic phenomena:
One usage refers to a variety (linguisti ...
s, a trait that is rare in the animal kingdom.[
]
Tool use
Elephants show a remarkable ability to use tools
A tool is an object that can extend an individual's ability to modify features of the surrounding environment or help them accomplish a particular task. Although many animals use simple tools, only human beings, whose use of stone tools dates ba ...
, using their trunks like arms. Elephants have been observed digging holes to drink water and then ripping bark from a tree, chewing it into the shape of a ball, filling in the hole and covering over it with sand to avoid evaporation
Evaporation is a type of vaporization that occurs on the surface of a liquid as it changes into the gas phase. High concentration of the evaporating substance in the surrounding gas significantly slows down evaporation, such as when humidi ...
, then later going back to drink from the same spot. They also often use branches to swat flies or scratch themselves. Asian elephants have also been known to drop large rocks onto an electric fence to break the fence or to cut off the electricity.[ Asian elephants in India have been known to break electric fences using logs and clear the surrounding wires using their tusks to provide a safe passageway.
]
Art and music
Like several other species that are able to produce abstract art, elephants, using their trunks to hold brushes, create paintings which some have compared to the work of abstract expressionists. Elephant art is now commonly featured at zoos, and is shown in museums and galleries around the world. Ruby
A ruby is a pinkish red to blood-red colored gemstone, a variety of the mineral corundum ( aluminium oxide). Ruby is one of the most popular traditional jewelry gems and is very durable. Other varieties of gem-quality corundum are called sa ...
at the Phoenix Zoo
The Phoenix Zoo opened in 1962 and is the largest privately owned, non-profit zoo in the United States. Located in Phoenix, Arizona, the zoo was founded by Robert Maytag, a member of the Maytag family, and operates on of land in the Papago Park ...
is considered the original elephant art star, and her paintings have sold for as much as $25,000. Ruby chose her own colors and was said to have a keen sense of which color she wished to use. The Asian Elephant Art & Conservation Project, an "elephant art academy" in New York, teaches retired elephants to paint. For paintings that resemble identifiable objects, teachers give the elephants guidance. An example of this was shown in the TV program ''Extraordinary Animals'', in which elephants at a camp in Thailand were able to draw portraits with flowers. Although the images were drawn by the elephants, there was always a trainer assisting and guiding the movement.
A popular video showing an elephant painting a picture of another elephant became widespread on Internet news and video websites. The website Snopes.com, which specializes in debunking urban legends, lists the video as "partly true", in that the elephant produced the brush strokes, but notes that the similarity of the produced paintings is indicative of a learned sequence of strokes rather than a creative effort on the part of the elephant.
It was noted by ancient Romans and Asian elephant handlers (mahout
A mahout is an elephant rider, trainer, or keeper. Mahouts were used since antiquity for both civilian and military use. Traditionally, mahouts came from ethnic groups with generations of elephant keeping experience, with a mahout retaining h ...
s) that elephants can distinguish melodies. Performing circus elephants commonly follow musical cues, and Adam Forepaugh
Adam John Forepaugh (born Adam John Forbach; February 28, 1831 – January 22, 1890) was an American horse trader and circus owner. From 1865 through 1890 his circus operated under various names including Forepaugh's Circus, Forepaugh's Gigantic ...
and Barnum & Bailey circuses even featured "elephant bands". German evolutionary biologist Bernhard Rensch
Bernhard Rensch (21 January 1900 – 4 April 1990) was a German evolutionary biologist and ornithologist who did field work in Indonesia and India. Starting his scientific career with pro-Lamarckian views, he shifted to selectionism and became ...
studied an elephant's ability to distinguish music, and in 1957 published the results in ''Scientific American''. Rensch's test elephant could distinguish 12 tones in the music scale and could remember simple melodies. Even though played on varying instruments and at different pitches, timbres and meters, she recognized the tones a year and a half later. These results have been backed up by the Human-Elephant Learning Project which studies elephant intelligence.
An elephant named Shanthi at the National Zoo in Washington D.C. displayed the ability to play the harmonica
The harmonica, also known as a French harp or mouth organ, is a free reed wind instrument used worldwide in many musical genres, notably in blues, American folk music, classical music, jazz, country, and rock. The many types of harmonica inclu ...
and various horn instrument
A horn is any of a family of musical instruments made of a tube, usually made of metal and often curved in various ways, with one narrow end into which the musician blows, and a wide end from which sound emerges. In horns, unlike some other br ...
s. She reportedly always ended her songs with a crescendo
In music, the dynamics of a piece is the variation in loudness between notes or phrases. Dynamics are indicated by specific musical notation, often in some detail. However, dynamics markings still require interpretation by the performer dependi ...
.
Recording group Thai Elephant Orchestra
The Thai Elephant Orchestra is a musical ensemble consisting of as many as fourteen Thai elephants near Lampang in Northern Thailand. The elephants play music, essentially as conducted improvisations, on specially designed heavy-duty musical ins ...
is an ensemble of elephants who improvise music on specially made instruments with minimal interaction from their handlers. The orchestra was co-founded by pachyderm expert Richard Lair, who works at the Thai Elephant Conservation Center in Lampang, and David Sulzer (artist name, Dave Soldier
David Sulzer (born November 6, 1956) is an American neuroscientist and musician. He is a professor at Columbia University Medical Center in the departments of psychiatry, neurology, and pharmacology. Sulzer's laboratory investigates the interact ...
) who studies the role of dopaminergic
Dopaminergic means "related to dopamine" (literally, "working on dopamine"), dopamine being a common neurotransmitter. Dopaminergic substances or actions increase dopamine-related activity in the brain. Dopaminergic brain pathways facilitate d ...
synapse
In the nervous system, a synapse is a structure that permits a neuron (or nerve cell) to pass an electrical or chemical signal to another neuron or to the target effector cell.
Synapses are essential to the transmission of nervous impulses from ...
s in memory consolidation
Memory consolidation is a category of processes that stabilize a memory trace after its initial acquisition. A memory trace is a change in the nervous system caused by memorizing something. Consolidation is distinguished into two specific processe ...
, learning, and behavior at Columbia University
Columbia University (also known as Columbia, and officially as Columbia University in the City of New York) is a private research university in New York City. Established in 1754 as King's College on the grounds of Trinity Church in Manhatt ...
. According to neurobiologist Aniruddh Patel, the orchestra's star drummer named Pratidah, exhibits musicality, stating: "Either when drumming alone or with the orchestra, Pratidah was remarkably steady". He also noted that she developed a swing-type rhythm pattern when playing with other elephants.
Problem-solving ability
Elephants are able to spend substantial time working on problems. They are able to change their behavior radically to face new challenges, a hallmark of complex intelligence
Intelligence has been defined in many ways: the capacity for abstraction, logic, understanding, self-awareness, learning, emotional knowledge, reasoning, planning, creativity, critical thinking, and problem-solving. More generally, it can b ...
.
Problem-solving experiments
A 2010 experiment revealed that in order to reach food, "elephants can learn to coordinate with a partner in a task requiring two individuals to simultaneously pull two ends of the same rope to obtain a reward", putting them on an equal footing with chimpanzees in terms of their level of cooperative skills.
A study by Dr. Naoko Irie of Tokyo University has shown that elephants demonstrate skills at arithmetic. The experiment "consist dof dropping varying numbers of apples into two buckets in front of the eno Zooelephants and then recording how often they could correctly choose the bucket holding the most fruit." When more than one apple was being dropped into the bucket, this meant that the elephants had to "keep running totals in their heads to keep track of the count." The results showed that "Seventy-four percent of the time, the animals correctly picked the fullest bucket. An African elephant named Ashya scored the highest with an amazing eighty-seven percent … Humans in this same contest managed a success rate of just sixty-seven percent." The study was also filmed to ensure its accuracy.
A study on Discovery News found that elephants, during an intelligence test employing food rewards, had found shortcuts that not even the experiment's researchers had thought of.
Adaptive behavior in captivity
In the 1970s, at Marine World Africa, USA, there lived an Asian elephant named Bandula.
Bandula worked out how to break open or unlock several of the pieces of equipment used to keep the shackles on her feet secure. The most complex device was a Brummel hook, a device that closes when two opposite points are slid together. Bandula used to fiddle with the hook until it slid apart when it was aligned. Once she had freed herself, she would help the other elephants escape.
In Bandula's case and certainly with other captive elephants, there was an element of deception
Deception or falsehood is an act or statement that misleads, hides the truth, or promotes a belief, concept, or idea that is not true. It is often done for personal gain or advantage. Deception can involve dissimulation, propaganda and sleight o ...
involved during the escapes, such as the animals
Animals are multicellular, eukaryotic organisms in the biological kingdom Animalia. With few exceptions, animals consume organic material, breathe oxygen, are able to move, can reproduce sexually, and go through an ontogenetic stage in ...
looking around making sure no one was watching.[
In another case, a female elephant worked out how she could unscrew iron rods with an eye hole that was an inch (2.5 cm) thick. She used her trunk to create leverage and then untwisted the bolt.][
]Ruby
A ruby is a pinkish red to blood-red colored gemstone, a variety of the mineral corundum ( aluminium oxide). Ruby is one of the most popular traditional jewelry gems and is very durable. Other varieties of gem-quality corundum are called sa ...
, an Asian elephant at Phoenix Zoo
The Phoenix Zoo opened in 1962 and is the largest privately owned, non-profit zoo in the United States. Located in Phoenix, Arizona, the zoo was founded by Robert Maytag, a member of the Maytag family, and operates on of land in the Papago Park ...
would often eavesdrop on conversations keepers would have talking about her. When she heard the word ''paint'', she became very excitable. The colors she favored were green, yellow, blue and red. Once, a fire truck came and parked outside her enclosure where a man had just had a heart attack. The lights on the truck were flashing red, white and yellow. When Ruby painted later on in the day, she chose those colors. She also showed a preference for colors that the keepers wore.[
Harry Peachey, an elephant trainer, developed a cooperative relationship with an elephant named Koko. Koko would help the keepers out, "prompting" them to encourage him with various commands and words that Koko would learn. Peachey stated that elephants are almost predisposed to cooperate and work with humans as long as they are treated with respect and sensitivity. Koko worked out when his keepers needed a bit of "elephant help" when they were transferring the females of the group to another zoo. When the keepers wanted to transfer a female, they would usually say her name, followed by the word ''transfer'' (e.g., "Connie transfer"). Koko soon figured out what this meant. If the keepers asked an elephant to transfer and it did not budge, they would say, "Koko, give me a hand." When he heard this, Koko would help. After 27 years of working with elephants, Peachey firmly believes that they can understand the ]semantics
Semantics (from grc, σημαντικός ''sēmantikós'', "significant") is the study of reference, meaning, or truth. The term can be used to refer to subfields of several distinct disciplines, including philosophy
Philosophy (f ...
and syntax
In linguistics, syntax () is the study of how words and morphemes combine to form larger units such as phrases and sentences. Central concerns of syntax include word order, grammatical relations, hierarchical sentence structure ( constituency) ...
of some of the words they hear. This is something thought to be very rare in the animal kingdom.[
According to one source, elephants can figure out how to retrieve distant objects that they cannot otherwise reach by using a stick.
]
Adaptive behavior in the wild
In the wild, elephants display clever methods of finding resources. Elephants have keen memories, and when evaluating foraging
Foraging is searching for wild food resources. It affects an animal's Fitness (biology), fitness because it plays an important role in an animal's ability to survive and reproduce. Optimal foraging theory, Foraging theory is a branch of behaviora ...
locations, respond more strongly to long‐term patterns of productivity
Productivity is the efficiency of production of goods or services expressed by some measure. Measurements of productivity are often expressed as a ratio of an aggregate output to a single input or an aggregate input used in a production proces ...
than to immediate forage conditions. In times of scarcity, they return to areas which have been reliable over many years rather than the last sites visited. They also favor travelling on dirt roads in the dry season, as easy walking terrain to conserve energy.
Although it is common for herbivores to find salt licks
A mineral lick (also known as a salt lick) is a place where animals can go to lick essential mineral nutrients from a deposit of salts and other minerals. Mineral licks can be naturally occurring or artificial (such as blocks of salt that farm ...
or to ingest inorganic matter for sodium, elephants in the Mount Elgon National Park, Kenya, have learned to venture deep into Kitum Cave
Kitum Cave is located in Mount Elgon National Park, Kenya. In the 1980s, two European visitors contracted Marburg virus disease there. It is one of five named “elephant caves” of Mount Elgon where animals, including elephants, “mine” the ...
to utilize its minerals in what has been described as 'quarrying' and 'salt mining'. Although the elephants clearly do not understand that they require salt in their diet, they show interest only in the cation
An ion () is an atom or molecule with a net electrical charge.
The charge of an electron is considered to be negative by convention and this charge is equal and opposite to the charge of a proton, which is considered to be positive by convent ...
-rich zeolite
Zeolites are microporous, crystalline aluminosilicate materials commonly used as commercial adsorbents and catalysts. They mainly consist of silicon, aluminium, oxygen, and have the general formula ・y where is either a metal ion or H+. These pos ...
, tusking it into smaller edible fragments. This activity is performed in groups, and years of tusk marks indicate the knowledge of the cave has been passed down over generations. Poaching
Poaching has been defined as the illegal hunting or capturing of wild animals, usually associated with land use rights.
Poaching was once performed by impoverished peasants for subsistence purposes and to supplement meager diets. It was set a ...
has caused the elephants to alter their behavior and avoid the more widely known caves.
Applying the string-drawing task to elephants
In 1956, W. H. Thorpe explained: The ability to pull up food which is suspended by a thread, the pulled in loop being held by the foot while the bird reaches with its beak for the next pull, is doubtfully inborn and it has been subject to many experiments. The act appears at first sight to be a real and sudden solution of the problem from the start, and thus to qualify for inclusion under 'insight learning.' Successful performance in this task has been documented in well over ten bird species.
More recently, Bernd Heinrich
Bernd Heinrich (born April 19, 1940 in Bad Polzin, Germany), is a professor emeritus in the biology department at the University of Vermont and is the author of a number of books about nature writing and biology. Heinrich has made major contribut ...
and Thomas Bugnyar concluded that ravens' "behaviour in accessing meat on a string is not only a product of rapid learning but may involve some understanding of cause–effect relation between string, food and certain body parts."
String-pulling behavior has been likewise studied in seven Asian elephants by presenting them with a retractable (bungee) cord. In this setup, the cord is tied to a heavy log a few meters away from the elephant. A sugarcane (a favorite elephant treat) is attached to the cord, and can only be retrieved by repeated, coordinated, action of the trunk and another body part. The results were clearcut:All seven logging elephants fully mastered the string-drawing sequence within 1–3 experimental sessions. In all cases of retractable rope pulling, the sequence involved pulling by the trunk, and then securing the rope by either foot or mouth. After 2–6 coordinated pulls, while still holding the rope with either mouth or foot, the elephants disengaged the sugarcane from the rope while still using mouth or forefoot as an anchor, and then consumed the sugarcane. All elephants seemed to be flexible about the use of anchor, interchangeably using mouth, foreleg, or both.
Self-awareness
Elephants have joined a small group of animals, including great apes
The Hominidae (), whose members are known as the great apes or hominids (), are a taxonomic family of primates that includes eight extant species in four genera: '' Pongo'' (the Bornean, Sumatran and Tapanuli orangutan); ''Gorilla'' (the east ...
, bottlenose dolphins
Bottlenose dolphins are aquatic mammals in the genus ''Tursiops.'' They are common, cosmopolitan members of the family Delphinidae, the family of oceanic dolphins. Molecular studies show the genus definitively contains two species: the comm ...
and Eurasian magpie
The Eurasian magpie or common magpie (''Pica pica'') is a resident breeding bird throughout the northern part of the Eurasian continent. It is one of several birds in the crow family (corvids) designated magpies, and belongs to the Holarctic ra ...
s, that exhibit self-awareness
In philosophy of self, self-awareness is the experience of one's own personality or individuality. It is not to be confused with consciousness in the sense of qualia. While consciousness is being aware of one's environment and body and lifesty ...
. The study was conducted with the Wildlife Conservation Society (WCS) using elephants at the Bronx Zoo
The Bronx Zoo (also historically the Bronx Zoological Park and the Bronx Zoological Gardens) is a zoo within Bronx Park in the Bronx, New York. It is one of the largest zoos in the United States by area and is the largest metropolitan zoo in ...
in New York. Although many animals respond to a mirror, very few show any evidence that they recognize it is in fact themselves in the mirror reflection.
The Asian elephants in the study also displayed this type of behavior when standing in front of a mirror – they inspected the mirror and brought food close to the mirror for consumption.
Evidence of elephant self-awareness was shown when the elephant Happy repeatedly touched a painted ''X'' on her head with her trunk, a mark which could only be seen in the mirror. Happy ignored another mark made with colorless paint that was also on her forehead to ensure she was not merely reacting to a smell or feeling.
Frans De Waal, who ran the study, stated, "These parallels between humans and elephants suggest a convergent cognitive evolution possibly related to complex society and cooperation."
Self-awareness and culling
There has been considerable debate over the issue of culling
In biology, culling is the process of segregating organisms from a group according to desired or undesired characteristics. In animal breeding, it is the process of removing or segregating animals from a breeding stock based on a specific tr ...
African elephants in South Africa's Kruger National Park
Kruger National Park is a South African National Park and one of the largest game reserves in Africa. It covers an area of in the provinces of Limpopo and Mpumalanga in northeastern South Africa, and extends from north to south and from ea ...
as a means of controlling the population. Some scientists and environmentalists argue that it is "unnecessary and inhumane" to cull them since "elephants resemble humans in a number of ways, not least by having massive brains, social bonds that appear to be empathetic, long gestations, high intelligence, offspring that require an extended period of dependent care, and long life spans." A South African animal rights group asked in a statement anticipating the announcement, "How much like us do elephants have to be before killing them becomes murder?"
Others argue that culling is necessary when biodiversity is threatened. However, the protection of biodiversity argument has been questioned by some animal rights advocates who argue that the animal which most greatly threatens and damages biodiversity is humanity, so if we are not willing to cull our own species we cannot morally justify culling another.
Arguments against intelligence
This section summarizes experiments that cannot be readily reconciled with the view that elephants are highly intelligent. These experiments, in turn, rely on pioneering early work with dogs and cats.
Discrimination tasks
Edward Thorndike
Edward Lee Thorndike (August 31, 1874 – August 9, 1949) was an American psychologist who spent nearly his entire career at Teachers College, Columbia University. His work on comparative psychology and the learning process led to the theory o ...
argued that his cats and dogs escaped puzzle boxes through a mindless process of trial and error. Because understanding something as simple as pulling a loop to open a door must occur rapidly or not at all, it should have induced, at some point during the repeated introductions of his animals into the box, a sudden reduction in escape time. The actual, gradual, slope of the time-curve that he did observe suggested to him that his subjects failed to understand the cause-effect relationships between their actions and escape.
In 1957, researchers reported that a young Asian elephant needed 330 trials, over a period of several days, to consistently choose the reinforced response in her first discrimination task. In an experiment which employed another sense modality, an 8-year-old took 7.5 months to distinguish 12 tones.
Similarly, in discrimination experiments with Asian elephants in the logging camps of Myanmar, only 13 Burmese elephants mastered black/white or large/small visual discrimination tasks, while 7 elephants failed to master the task.
Like Thorndike's cats and dogs, the 13 elephants that did master a black/white or large/small discrimination task did so gradually, over several sessions. The slope of the time-curve again suggested the elephants failed to understand the cause-effect relationships between lid removal and food retrieval.
The pre-training that preceded these Myanmar discrimination experiments involved learning to remove a lid from a bucket or to displace a box to uncover a hole in the ground. On average, the 20 elephants required 3.4 sessions to master the pre-training task.
Possible absence of causal reasoning
Thorndike's dogs and cats learned to escape a box by, for instance, pulling a loop attached to a string that opened the box. On subsequent introductions to an open box when string-pulling no longer served a purpose, the animal continued to pull a string before getting out of the box. Thorndike concluded that the animal solved the task mechanically, without understanding the causal link between string-pulling and escaping.
A conceptually similar experiment involved pre-training four logging Asian elephants to remove food from a cover-less bucket by inserting their trunk into the bucket. Next, a treat was placed at the bottom of a bucket and, at the same time, the bucket was covered with a lid. The elephants were then trained to obtain the treat by removing the lid. Once this behavioral sequence was established, a treat was placed inside the bucket and, at the same time, the lid was placed on the ground alongside the bucket, so that the lid no longer obstructed access to the food.
As in the Thorndike case, if the elephants understand the nature of the task, they should ignore the lid on the ground and retrieve the reward directly, as they used to do in pre-training sessions before the lid was introduced. But if they fail to understand the causal link between lid removal and food retrieval, they might continue to remove the lid before retrieving the food. The observations supported Thorndike's mechanical learning hypothesis.
See also
* Animal cognition
Animal cognition encompasses the mental capacities of non-human animals including insect cognition. The study of animal conditioning and learning used in this field was developed from comparative psychology. It has also been strongly influenc ...
* Spindle neuron
Von Economo neurons (VENs), also called spindle neurons, are a specific class of mammalian cortical neurons characterized by a large spindle-shaped soma (or body) gradually tapering into a single apical axon (the ramification that ''transmits ...
* Tool use
Tool use by animals is a phenomenon in which an animal uses any kind of tool in order to achieve a goal such as acquiring food and water, grooming, defence, communication, recreation or construction. Originally thought to be a skill possessed ...
* Vocal learning Vocal learning is the ability to modify acoustic and syntactic sounds, acquire new sounds via imitation, and produce vocalizations. "Vocalizations" in this case refers only to sounds generated by the vocal organ (mammalian larynx or avian syrinx) as ...
References
External links
The Truth Behind Elephant Brainpower
by BBC News
BBC News is an operational business division of the British Broadcasting Corporation (BBC) responsible for the gathering and broadcasting of news and current affairs in the UK and around the world. The department is the world's largest broadca ...
Elephant Cognition in Primate Perspective
by Richard W. Byrne and Lucy A. Bates
Do Elephants Show Empathy?
by the ''Journal of Consciousness Studies''
African Elephants have Expectations about the Locations of out-of-sight Family Members
by the Amboseli Trust for Elephants
{{DEFAULTSORT:Elephant Intelligence
Animal intelligence
Elephants
Articles containing video clips