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The Elateroidea are a large superfamily of
beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
s. It contains the familiar click beetles, fireflies, and soldier beetles and their relatives. It consists of about 25,000 species.


Description

Elateroidea is a morphologically diverse group, including hard-bodied beetles with 5 abdominal ventrites, soft-bodied beetles with 7-8 ventrites connected with membranes (formerly known as cantharoids), and beetles with intermediate forms. They have a range of sizes and colours, but in terms of shape, they are usually narrow and parallel-sided as adults. Many of the sclerotised elateroids ( Cerophytidae,
Eucnemidae Eucnemidae, or false click beetles, are a family of elateroid beetles including about 1700 species distributed worldwide. Description Closely related to the family Elateridae, specimens of Eucnemidae can reach a length of . Bodies are slight ...
, Throscidae, Elateridae) have a clicking mechanism. This is a peg on the prothorax which fits into a cavity in the mesothorax. When a click beetle bends its body, the peg snaps into the cavity, causing the beetle's body to straighten so suddenly that it jumps into the air. Most beetles capable of bioluminescence are in the Elateroidea, in the families Lampyridae (~2000 species), Phengodidae (~200 species), Rhagophthalmidae (100 species) and Elateridae (>100 species). Females in several lineages, including
Lycidae The Lycidae are a family in the beetle order Coleoptera, members of which are commonly called net-winged beetles. These beetles are cosmopolitan, being found in Nearctic, Palearctic, Neotropical, Afrotropical, Oriental, and Australian ecoregions ...
, Lampyridae, Phengogidae and Rhagophthalmidae, do not pupate and remain in a larval form. This trait is estimated to have evolved independently at least three times within the superfamily. Some Elateroidea, including species of Cantharidae and Lycidae, have bright aposematic colours to signal to predators that they are poisonous and so should not be eaten.


Families

The validity and relationships of some families, such as
Podabrocephalidae ''Podabrocephalus'' is an enigmatic monotypic genus in the family Ptilodactylidae. For decades it had historically been placed in its own family, Podabrocephalidae, or sometimes included in the highly diverse family Cerambycidae The longhorn be ...
are not fully resolved. The family
Rhinorhipidae ''Rhinorhipus'' is a genus of beetles that contains a single species, ''Rhinorhipus tamborinensis'' from southern Queensland, Australia. It is the sole member of the family Rhinorhipidae and superfamily Rhinorhipoidea. It is an isolated lineage n ...
has recently been removed to its own superfamily, with evidence that it is a basal taxon within Elateriformia dating to an Upper Triassic/Lower Jurassic split from other extant beetle lineages. * Artematopodidae Lacordaire, 1857 – soft-bodied plant beetles (= Eurypogonidae) * Brachypsectridae Leconte & Horn, 1883 – Texas beetles * Cantharidae Imhoff, 1856 – soldier beetles * Cerophytidae Latreille, 1834 – rare click beetles * Elateridae Leach, 1815 – click beetles (including Ampedidae, Balgidae, Dicronychidae, Drilidae, Lissomidae, Plastoceridae, Prosternidae, Protelateridae, Pyrophoridae, Synaptidae) *
Eucnemidae Eucnemidae, or false click beetles, are a family of elateroid beetles including about 1700 species distributed worldwide. Description Closely related to the family Elateridae, specimens of Eucnemidae can reach a length of . Bodies are slight ...
Eschscholtz, 1829false click beetles (including Anischiidae and Perothopidae) * Iberobaeniidae Bocak ''et al.'', 2016 * Jurasaidae Rosa, Costa, Klamp & Kundrata, 2020 *
Lycidae The Lycidae are a family in the beetle order Coleoptera, members of which are commonly called net-winged beetles. These beetles are cosmopolitan, being found in Nearctic, Palearctic, Neotropical, Afrotropical, Oriental, and Australian ecoregions ...
– net-winged beetles * Omethidae LeConte, 1861false firefly beetles (including Telegeusidae) * Throscidae Laporte, 1840false metallic wood-boring beetles (= Trixagidae) * † Mysteriomorphidae Alekseev and Ellenberger, 2019 (Cretaceous) * Lampyroid clade ** † Cretophengodidae Li, Kundrata, Tihelka & Cai, 2021 (Cretaceous) ** Lampyridae Latreille, 1817 – firefly beetles ** Phengodidae LeConte 1861 – glowworm beetles (including Cydistinae) ** Rhagophthalmidae Olivier, 1907 (sometimes in Phengodidae, might belong in Lampyridae) ** Sinopyrophoridae Bi & Li, 2019 *''Incertae sedis'': **†'' Anoeuma'' Li, Kundrata & Cai, 2022 (Cretaceous)


Phylogeny

Some morphological and molecular phylogenetic analyses find that Byrrhoidea is either a monophyletic or paraphyletic group closely related to Elateroidea. Based on Kusy et al. 2018 and 2020


References


External links


Tree of Life - Elateroidea
{{DEFAULTSORT:Elateroidea Beetle superfamilies