Eifelian Life
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The Eifelian is the first of two
faunal stage In chronostratigraphy, a stage is a succession of rock strata laid down in a single age on the geologic timescale, which usually represents millions of years of deposition. A given stage of rock and the corresponding age of time will by conventi ...
s in the
Middle Devonian The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, wher ...
Epoch. It lasted from 393.3 ± 1.2 million years ago to 387.7 ± 0.8 million years ago. It was preceded by the
Emsian The Emsian is one of three faunal stages in the Early Devonian Epoch. It lasted from 407.6 ± 2.6 million years ago to 393.3 ± 1.2 million years ago. It was preceded by the Pragian Stage and followed by the Eifelian Stage. It is named after the ...
Stage and followed by the
Givetian The Givetian is one of two faunal stages in the Middle Devonian Period. It lasted from million years ago to million years ago. It was preceded by the Eifelian Stage and followed by the Frasnian Stage. It is named after the town of Givet in Fra ...
Stage. North American subdivisions of the Eifelian Stage include Southwood, and part of Cazenovia (or Cazenovian).


Name and definition

The Eifelian is named after the
Eifel Mountains The Eifel (; lb, Äifel, ) is a low mountain range in western Germany and eastern Belgium. It occupies parts of southwestern North Rhine-Westphalia, northwestern Rhineland-Palatinate and the southern area of the German-speaking Community of ...
of Western
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
, which exposed the
GSSP A Global Boundary Stratotype Section and Point (GSSP) is an internationally agreed upon reference point on a stratigraphic section which defines the lower boundary of a stage on the geologic time scale. The effort to define GSSPs is conducted b ...
section at the Wetteldorf Richtschnitt outcrop. The base of the Eifelian is defined by the start of the '' Polygnathus partitus''
conodont Conodonts (Greek ''kōnos'', "cone", + ''odont'', "tooth") are an extinct group of agnathan (jawless) vertebrates resembling eels, classified in the class Conodonta. For many years, they were known only from their tooth-like oral elements, which ...
zone. This layer lies within the Upper Heisdorf Formation, below the base of the Lauch Formation.


Extinctions

The end of the Eifelian was marked by a biological crisis known as the Kačák Event, a two-part interval of extinction which led to ecological turnover among
ammonoids Ammonoids are a group of extinct marine mollusc animals in the subclass Ammonoidea of the class Cephalopoda. These molluscs, commonly referred to as ammonites, are more closely related to living coleoids (i.e., octopuses, squid and cuttle ...
,
conodonts Conodonts (Greek ''kōnos'', "cone", + ''odont'', "tooth") are an extinct group of agnathan (jawless) vertebrates resembling eels, classified in the class Conodonta. For many years, they were known only from their tooth-like oral elements, which ...
, and other free-swimming animals. In deep marine waters, the event is indicated by
anoxic The term anoxia means a total depletion in the level of oxygen, an extreme form of hypoxia or "low oxygen". The terms anoxia and hypoxia are used in various contexts: * Anoxic waters, sea water, fresh water or groundwater that are depleted of diss ...
black shales. There is evidence for a major pulse of transgression (
sea level rise Globally, sea levels are rising due to human-caused climate change. Between 1901 and 2018, the globally averaged sea level rose by , or 1–2 mm per year on average.IPCC, 2019Summary for Policymakers InIPCC Special Report on the Ocean and Cry ...
) and warming during the event. Warming and sea level rise through the Eifelian and beyond would have had major effects on diversity, likely leading to the downfall of several marine
biogeographic realms Biogeography is the study of the species distribution, distribution of species and ecosystems in geography, geographic space and through evolutionary history of life, geological time. Organisms and biological community (ecology), communities of ...
. The cool-water Malvinokaffric Realm (MKR), on the northwest edge of
Gondwana Gondwana () was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic (about 550 million years ago) and began to break up during the Jurassic period (about 180 million years ago). The final stages ...
, was decimated as rising temperatures eliminated suitably
temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout t ...
habitat. Fauna of the Eastern Americas Realm (EAR), which was restricted to a shallow basin in southwest Laurussia, were gradually replaced by aggressive cosmopolitan species of the Old World Realm (OWR), which invaded through a seaway cutting along a flooded continental arch on the western edge of Laurussia.


References

Middle Devonian {{geochronology-stub