Eifelian Extinctions
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The Eifelian is the first of two faunal stages in the Middle Devonian Epoch. It lasted from 393.3 ± 1.2 million years ago to 387.7 ± 0.8 million years ago. It was preceded by the Emsian Stage and followed by the Givetian Stage. North American subdivisions of the Eifelian Stage include Southwood, and part of Cazenovia (or Cazenovian).


Name and definition

The Eifelian is named after the
Eifel Mountains The Eifel (; lb, Äifel, ) is a low mountain range in western Germany and eastern Belgium. It occupies parts of southwestern North Rhine-Westphalia, northwestern Rhineland-Palatinate and the southern area of the German-speaking Community of ...
of Western Germany, which exposed the GSSP section at the Wetteldorf Richtschnitt outcrop. The base of the Eifelian is defined by the start of the '' Polygnathus partitus''
conodont Conodonts (Greek ''kōnos'', "cone", + ''odont'', "tooth") are an extinct group of agnathan (jawless) vertebrates resembling eels, classified in the class Conodonta. For many years, they were known only from their tooth-like oral elements, which ...
zone. This layer lies within the Upper
Heisdorf Formation Heisdorf () is a town in the commune of Steinsel, in central Luxembourg Luxembourg ( ; lb, Lëtzebuerg ; french: link=no, Luxembourg; german: link=no, Luxemburg), officially the Grand Duchy of Luxembourg, ; french: link=no, Grand-Duché ...
, below the base of the
Lauch Formation The Lauch Formation is a geologic formation in Germany. It preserves fossils dating back to the Devonian period. See also * List of fossiliferous stratigraphic units in Germany See also * Lists of fossiliferous stratigraphic units in Europ ...
.


Extinctions

The end of the Eifelian was marked by a biological crisis known as the Kačák Event, a two-part interval of extinction which led to ecological turnover among ammonoids, conodonts, and other free-swimming animals. In deep marine waters, the event is indicated by anoxic black shales. There is evidence for a major pulse of
transgression Transgression may refer to: Legal, religious and social *Sin, a violation of God's Ten Commandments or other elements of God's moral law *Crime, legal transgression, usually created by a violation of social or economic boundary **In civil law ju ...
( sea level rise) and warming during the event. Warming and sea level rise through the Eifelian and beyond would have had major effects on diversity, likely leading to the downfall of several marine
biogeographic realms Biogeography is the study of the species distribution, distribution of species and ecosystems in geography, geographic space and through evolutionary history of life, geological time. Organisms and biological community (ecology), communities of ...
. The cool-water Malvinokaffric Realm (MKR), on the northwest edge of
Gondwana Gondwana () was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic (about 550 million years ago) and began to break up during the Jurassic period (about 180 million years ago). The final stages ...
, was decimated as rising temperatures eliminated suitably temperate habitat. Fauna of the Eastern Americas Realm (EAR), which was restricted to a shallow basin in southwest Laurussia, were gradually replaced by aggressive cosmopolitan species of the Old World Realm (OWR), which invaded through a seaway cutting along a flooded continental arch on the western edge of Laurussia.


References

Middle Devonian {{geochronology-stub