An ecoregion (ecological region) or ecozone (ecological zone) is an
ecologically
Ecology () is the study of the relationships between living organisms, including humans, and their biophysical environment, physical environment. Ecology considers organisms at the individual, population, community (ecology), community, ecosy ...
and geographically defined area that is smaller than a
bioregion
A bioregion is an ecologically and geographically defined area that is smaller than a biogeographic realm, but larger than an ecoregion or an ecosystem, in the World Wide Fund for Nature classification scheme. There is also an attempt to use the ...
, which in turn is smaller than a
biogeographic realm
A biogeographic realm or ecozone is the broadest biogeographic division of Earth's land surface, based on distributional patterns of terrestrial organisms. They are subdivided into bioregions, which are further subdivided into ecoregions.
De ...
. Ecoregions cover relatively large areas of land or water, and contain characteristic, geographically distinct assemblages of natural
communities and
species
In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
. The
biodiversity
Biodiversity or biological diversity is the variety and variability of life on Earth. Biodiversity is a measure of variation at the genetic (''genetic variability''), species (''species diversity''), and ecosystem (''ecosystem diversity'') l ...
of
flora
Flora is all the plant life present in a particular region or time, generally the naturally occurring (indigenous (ecology), indigenous) native plant, native plants. Sometimes bacteria and fungi are also referred to as flora, as in the terms '' ...
,
fauna
Fauna is all of the animal life present in a particular region or time. The corresponding term for plants is ''flora'', and for fungi, it is ''funga''. Flora, fauna, funga and other forms of life are collectively referred to as ''Biota (ecology ...
and
ecosystems
An ecosystem (or ecological system) consists of all the organisms and the physical environment with which they interact. These biotic and abiotic components are linked together through nutrient cycles and energy flows. Energy enters the syst ...
that characterise an ecoregion tends to be distinct from that of other ecoregions. In theory, biodiversity or conservation ecoregions are relatively large areas of land or water where the probability of encountering different species and communities at any given point remains relatively constant, within an acceptable range of variation (largely undefined at this point).
Three caveats are appropriate for all bio-geographic mapping approaches. Firstly, no single bio-geographic framework is optimal for all taxa. Ecoregions reflect the best compromise for as many taxa as possible. Secondly, ecoregion boundaries rarely form abrupt edges; rather,
ecotone
An ecotone is a transition area between two biological communities, where two communities meet and integrate. It may be narrow or wide, and it may be local (the zone between a field and forest) or regional (the transition between forest and gras ...
s and mosaic
habitat
In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
s bound them. Thirdly, most ecoregions contain habitats that differ from their assigned
biome
A biome () is a biogeographical unit consisting of a biological community that has formed in response to the physical environment in which they are found and a shared regional climate. Biomes may span more than one continent. Biome is a broader ...
. Biogeographic provinces may originate due to various barriers, including physical (plate tectonics, topographic highs), climatic (latitudinal variation, seasonal range) and ocean chemical related (salinity, oxygen levels).
History
The history of the term is somewhat vague, and it had been used in many contexts: forest classifications (Loucks, 1962),
biome
A biome () is a biogeographical unit consisting of a biological community that has formed in response to the physical environment in which they are found and a shared regional climate. Biomes may span more than one continent. Biome is a broader ...
classifications (Bailey, 1976, 2014),
biogeographic
Biogeography is the study of the distribution of species and ecosystems in geographic space and through geological time. Organisms and biological communities often vary in a regular fashion along geographic gradients of latitude, elevation, ...
classifications (
WWF/
Global 200
The Global 200 is the list of ecoregions identified by the World Wide Fund for Nature ( WWF), the global conservation organization, as priorities for conservation. According to WWF, an ecoregion is defined as a "relatively large unit of land or w ...
scheme of Olson & Dinerstein, 1998), etc.
[Bailey, R. G. 2014. ''Ecoregions: The Ecosystem Geography of the. Oceans and Continents''. 2nd ed., Springer, 180 pp.]
The concept of ecoregion applied by Bailey gives more importance to ecological criteria and climate, while the WWF concept gives more importance to biogeography, that is, the
Species distribution, distribution of distinct species assemblages.
Definition and categorization
An ecoregion is a "recurring pattern of ecosystems associated with characteristic combinations of
soil
Soil, also commonly referred to as earth or dirt
Dirt is an unclean matter, especially when in contact with a person's clothes, skin, or possessions. In such cases, they are said to become dirty.
Common types of dirt include:
* Debri ...
and
landform that characterise that region". Omernik (2004) elaborates on this by defining ecoregions as: "areas within which there is spatial coincidence in characteristics of
geographical
Geography (from Greek: , ''geographia''. Combination of Greek words ‘Geo’ (The Earth) and ‘Graphien’ (to describe), literally "earth description") is a field of science devoted to the study of the lands, features, inhabitants, and ...
phenomena associated with differences in the quality, health, and integrity of
ecosystems
An ecosystem (or ecological system) consists of all the organisms and the physical environment with which they interact. These biotic and abiotic components are linked together through nutrient cycles and energy flows. Energy enters the syst ...
". "Characteristics of geographical phenomena" may include
geology
Geology () is a branch of natural science concerned with Earth and other astronomical objects, the features or rocks of which it is composed, and the processes by which they change over time. Modern geology significantly overlaps all other Ea ...
,
physiography
Physical geography (also known as physiography) is one of the three main branches of geography. Physical geography is the branch of natural science which deals with the processes and patterns in the natural environment such as the atmosphere, h ...
, vegetation, climate,
hydrology
Hydrology () is the scientific study of the movement, distribution, and management of water on Earth and other planets, including the water cycle, water resources, and environmental watershed sustainability. A practitioner of hydrology is call ...
, terrestrial and
aquatic fauna
Fauna is all of the animal life present in a particular region or time. The corresponding term for plants is ''flora'', and for fungi, it is ''funga''. Flora, fauna, funga and other forms of life are collectively referred to as ''Biota (ecology ...
, and soils, and may or may not include the impacts of human activity (e.g. land use patterns, vegetation changes). There is significant, but not absolute, spatial correlation among these characteristics, making the delineation of ecoregions an imperfect science. Another complication is that environmental conditions across an ecoregion boundary may change very gradually, e.g. the prairie-forest transition in the midwestern United States, making it difficult to identify an exact dividing boundary. Such transition zones are called
ecotone
An ecotone is a transition area between two biological communities, where two communities meet and integrate. It may be narrow or wide, and it may be local (the zone between a field and forest) or regional (the transition between forest and gras ...
s.
Ecoregions can be categorized using an
algorithm
In mathematics and computer science, an algorithm () is a finite sequence of rigorous instructions, typically used to solve a class of specific problems or to perform a computation. Algorithms are used as specifications for performing ...
ic approach or a holistic, "weight-of-evidence" approach where the importance of various factors may vary. An example of the algorithmic approach is
Robert Bailey's work for the U.S. Forest Service, which uses a hierarchical classification that first divides land areas into very large regions based on climatic factors, and subdivides these regions, based first on dominant potential vegetation, and then by
geomorphology and soil characteristics. The weight-of-evidence approach is exemplified by James Omernik's work for the
United States Environmental Protection Agency
The Environmental Protection Agency (EPA) is an independent executive agency of the United States federal government tasked with environmental protection matters. President Richard Nixon proposed the establishment of EPA on July 9, 1970; it ...
, subsequently adopted (with modification) for
North America by the
Commission for Environmental Cooperation.
The intended purpose of ecoregion delineation may affect the method used. For example, the
WWF ecoregions were developed to aid in biodiversity conservation planning, and place a greater emphasis than the Omernik or Bailey systems on floral and faunal differences between regions. The WWF classification defines an ecoregion as:
A large area of land or water that contains a geographically distinct assemblage of natural communities that:
:(a) Share a large majority of their species and ecological dynamics;
:(b) Share similar environmental conditions, and;
:(c) Interact ecologically in ways that are critical for their long-term persistence.
According to WWF, the boundaries of an ecoregion approximate the original extent of the natural communities prior to any major recent disruptions or changes. WWF has identified 867 terrestrial ecoregions, and approximately 450 freshwater ecoregions across the Earth.
Importance
The use of the term ecoregion is an outgrowth of a surge of interest in ecosystems and their functioning. In particular, there is awareness of issues relating to spatial scale in the study and management of
landscapes
A landscape is the visible features of an area of land, its landforms, and how they integrate with natural or man-made features, often considered in terms of their aesthetic appeal.''New Oxford American Dictionary''. A landscape includes the p ...
. It is widely recognized that interlinked ecosystems combine to form a whole that is "greater than the sum of its parts". There are many attempts to respond to ecosystems in an integrated way to achieve "multi-functional" landscapes, and various interest groups from
agricultural researchers to
conservationists
The conservation movement, also known as nature conservation, is a political, environmental, and social movement that seeks to manage and protect natural resources, including animal, fungus, and plant species as well as their habitat for the f ...
are using the "ecoregion" as a unit of analysis.
The "
Global 200
The Global 200 is the list of ecoregions identified by the World Wide Fund for Nature ( WWF), the global conservation organization, as priorities for conservation. According to WWF, an ecoregion is defined as a "relatively large unit of land or w ...
" is the list of ecoregions identified by WWF as priorities for
conservation
Conservation is the preservation or efficient use of resources, or the conservation of various quantities under physical laws.
Conservation may also refer to:
Environment and natural resources
* Nature conservation, the protection and managem ...
.
Terrestrial
Terrestrial ecoregions are land ecoregions, as distinct from freshwater and marine ecoregions. In this context, ''
terrestrial
Terrestrial refers to things related to land or the planet Earth.
Terrestrial may also refer to:
* Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on or near the ground, as opposed to ...
'' is used to mean "of land" (soil and rock), rather than the more general sense "of
Earth
Earth is the third planet from the Sun and the only astronomical object known to harbor life. While large volumes of water can be found throughout the Solar System, only Earth sustains liquid surface water. About 71% of Earth's surfa ...
" (which includes land and oceans).
WWF (World Wildlife Fund)
ecologist
Ecology () is the study of the relationships between living organisms, including humans, and their physical environment. Ecology considers organisms at the individual, population, community, ecosystem, and biosphere level. Ecology overlaps wi ...
s currently divide the land surface of the Earth into eight
biogeographical realm
A biogeographic realm or ecozone is the broadest biogeographic division of Earth's land surface, based on distributional patterns of terrestrial organisms. They are subdivided into bioregions, which are further subdivided into ecoregions.
De ...
s containing 867 smaller terrestrial ecoregions (see
list
A ''list'' is any set of items in a row. List or lists may also refer to:
People
* List (surname)
Organizations
* List College, an undergraduate division of the Jewish Theological Seminary of America
* SC Germania List, German rugby unio ...
). The WWF effort is a synthesis of many previous efforts to define and classify ecoregions.
The eight realms follow the major floral and faunal boundaries, identified by botanists and zoologists, that separate the world's major plant and animal communities. Realm boundaries generally follow
continent
A continent is any of several large landmasses. Generally identified by convention rather than any strict criteria, up to seven geographical regions
In geography, regions, otherwise referred to as zones, lands or territories, are areas t ...
al boundaries, or major barriers to plant and animal distribution, like the
Himalayas
The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the planet's highest peaks, including the very highest, Mount Everest. Over 10 ...
and the
Sahara. The boundaries of ecoregions are often not as decisive or well recognized, and are subject to greater disagreement.
Ecoregions are classified by
biome
A biome () is a biogeographical unit consisting of a biological community that has formed in response to the physical environment in which they are found and a shared regional climate. Biomes may span more than one continent. Biome is a broader ...
type, which are the major global plant communities determined by rainfall and climate. Forests, grasslands (including savanna and shrubland), and deserts (including
xeric shrublands) are distinguished by climate (
tropical
The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in
the Southern Hemisphere at S. The tropics are also referred to ...
and
subtropical vs.
temperate
In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout ...
and
boreal climates) and, for forests, by whether the trees are predominantly
conifers (
gymnosperm
The gymnosperms ( lit. revealed seeds) are a group of seed-producing plants that includes conifers, cycads, '' Ginkgo'', and gnetophytes, forming the clade Gymnospermae. The term ''gymnosperm'' comes from the composite word in el, γυμν ...
s), or whether they are predominantly broadleaf (
Angiosperms) and mixed (broadleaf and conifer). Biome types like
Mediterranean forests, woodlands, and scrub;
tundra
In physical geography, tundra () is a type of biome where tree growth is hindered by frigid temperatures and short growing seasons. The term ''tundra'' comes through Russian (') from the Kildin Sámi word (') meaning "uplands", "treeless mou ...
; and
mangroves
A mangrove is a shrub or tree that grows in coastal saline or brackish water. The term is also used for tropical coastal vegetation consisting of such species. Mangroves are taxonomically diverse, as a result of convergent evolution in severa ...
host very distinct ecological communities, and are recognized as distinct biome types as well.
Marine
Marine ecoregion
A marine ecoregion is an ecoregion, or ecological region, of the oceans and seas identified and defined based on biogeographic characteristics.
Introduction
A more complete definition describes them as “Areas of relatively homogeneous species ...
s are: "Areas of relatively homogeneous species composition, clearly distinct from adjacent systems….In ecological terms, these are strongly cohesive units, sufficiently large to encompass ecological or life history processes for most sedentary species."
They have been defined by The Nature Conservancy (TNC) and World Wildlife Fund (WWF) to aid in
conservation
Conservation is the preservation or efficient use of resources, or the conservation of various quantities under physical laws.
Conservation may also refer to:
Environment and natural resources
* Nature conservation, the protection and managem ...
activities for
marine ecosystems
Marine ecosystems are the largest of Earth's aquatic ecosystems and exist in waters that have a high salt content. These systems contrast with freshwater ecosystems, which have a lower salt content. Marine waters cover more than 70% of the su ...
. Forty-three priority marine ecoregions were delineated as part of WWF's Global 200 efforts. The scheme used to designate and classify marine ecoregions is analogous to that used for terrestrial ecoregions. Major habitat types are identified: polar, temperate shelves and seas, temperate upwelling, tropical upwelling, tropical coral, pelagic (trades and westerlies), abyssal, and hadal (ocean trench). These correspond to the terrestrial
biomes.
The Global 200 classification of marine ecoregions is not developed to the same level of detail and comprehensiveness as that of the terrestrial ecoregions; only the priority conservation areas are listed.
See Global 200 Marine ecoregions for a full list of marine ecoregions.
In 2007, TNC and WWF refined and expanded this scheme to provide a system of comprehensive near shore (to 200 meters depth) Marine Ecoregions of the World (MEOW).
The 232 individual marine ecoregions are grouped into 62
marine provinces, which in turn group into 12
marine realms, which represent the broad latitudinal divisions of polar, temperate, and tropical seas, with subdivisions based on ocean basins (except for the southern hemisphere temperate oceans, which are based on continents).
Major marine biogeographic realms, analogous to the eight terrestrial
biogeographic realm
A biogeographic realm or ecozone is the broadest biogeographic division of Earth's land surface, based on distributional patterns of terrestrial organisms. They are subdivided into bioregions, which are further subdivided into ecoregions.
De ...
s, represent large regions of the ocean basins:
Arctic
The Arctic ( or ) is a polar region located at the northernmost part of Earth. The Arctic consists of the Arctic Ocean, adjacent seas, and parts of Canada (Yukon, Northwest Territories, Nunavut), Danish Realm (Greenland), Finland, Iceland, N ...
,
Temperate Northern Atlantic
The Temperate Northern Atlantic is a biogeographic region of the Earth's seas, comprising the temperate and subtropical waters of the North Atlantic Ocean and connecting seas, including the Mediterranean Sea, Black Sea, and northern Gulf of Mexic ...
,
Temperate Northern Pacific,
Tropical Atlantic
The Tropical Atlantic realm is one of twelve marine realms that cover the world's coastal seas and continental shelves.
The Tropical Atlantic covers both sides of the Atlantic. In the western Atlantic, it extends from Bermuda, southern Florida, ...
,
Western Indo-Pacific
The Western Indo-Pacific is a biogeographic region of the Earth's seas, comprising the tropical waters of the eastern and central Indian Ocean. It is part of the larger Indo-Pacific, which includes the tropical Indian Ocean, the western and centra ...
,
Central Indo-Pacific,
Eastern Indo-Pacific
The Eastern Indo-Pacific is a biogeographic region of the Earth's seas, comprising the tropical waters around island groups in the central Pacific Ocean. It includes most of Polynesia, except for New Zealand and the Kermadec Islands. It also inclu ...
,
Tropical Eastern Pacific
The Tropical Eastern Pacific is one of the twelve marine realms that cover the coastal waters and continental shelves of the world's oceans. The Tropical Eastern Pacific extends along the Pacific Coast of the Americas, from the southern tip of t ...
,
Temperate South America
Temperate South America is a biogeographic region of the Earth's seas, comprising the temperate and subtropical waters of South America, including both the Pacific and Atlantic coasts of the continent and adjacent islands. It also includes the rem ...
,
Temperate Southern Africa
Temperate Southern Africa is a biogeographic region of the Earth's seas, comprising the temperate waters of southern Africa, where the Atlantic Ocean and Indian Ocean meet. It includes the coast of South Africa and Namibia, and reaches into south ...
,
Temperate Australasia
Temperate Australasia is a biogeographic region of the Earth's seas, comprising the temperate and subtropical waters of Australia and New Zealand, including both the Indian Ocean and Pacific coasts of the continent and adjacent islands.
Tempera ...
, and
Southern Ocean
The Southern Ocean, also known as the Antarctic Ocean, comprises the southernmost waters of the World Ocean, generally taken to be south of 60° S latitude and encircling Antarctica. With a size of , it is regarded as the second-small ...
.
A similar system of identifying areas of the oceans for conservation purposes is the system of large marine ecosystems (LMEs), developed by the US
National Oceanic and Atmospheric Administration
The National Oceanic and Atmospheric Administration (abbreviated as NOAA ) is an United States scientific and regulatory agency within the United States Department of Commerce that forecasts weather, monitors oceanic and atmospheric conditio ...
(NOAA).
Freshwater
A freshwater ecoregion is a large area encompassing one or more freshwater systems that contains a distinct assemblage of natural freshwater communities and species. The freshwater species, dynamics, and environmental conditions within a given ecoregion are more similar to each other than to those of surrounding ecoregions and together form a conservation unit. Freshwater systems include
river
A river is a natural flowing watercourse, usually freshwater, flowing towards an ocean, sea, lake or another river. In some cases, a river flows into the ground and becomes dry at the end of its course without reaching another body of w ...
s,
streams,
lake
A lake is an area filled with water, localized in a basin, surrounded by land, and distinct from any river or other outlet that serves to feed or drain the lake. Lakes lie on land and are not part of the ocean, although, like the much large ...
s, and
wetland
A wetland is a distinct ecosystem that is flooded or saturated by water, either permanently (for years or decades) or seasonally (for weeks or months). Flooding results in oxygen-free (anoxic) processes prevailing, especially in the soils. The p ...
s. Freshwater ecoregions are distinct from terrestrial ecoregions, which identify biotic communities of the land, and marine ecoregions, which are biotic communities of the oceans.
A map of Freshwater Ecoregions of the World, released in 2008, has 426 ecoregions covering virtually the entire non-marine surface of the earth.
World Wildlife Fund
The World Wide Fund for Nature Inc. (WWF) is an international non-governmental organization founded in 1961 that works in the field of wilderness preservation and the reduction of human impact on the environment. It was formerly named the Wo ...
(WWF) identifies twelve major habitat types of freshwater ecoregions: Large lakes, large river deltas, polar freshwaters, montane freshwaters, temperate coastal rivers, temperate floodplain rivers and wetlands, temperate upland rivers, tropical and subtropical coastal rivers, tropical and subtropical floodplain rivers and wetlands, tropical and subtropical upland rivers, xeric freshwaters and endorheic basins, and oceanic islands. The freshwater major habitat types reflect groupings of ecoregions with similar biological, chemical, and physical characteristics and are roughly equivalent to biomes for terrestrial systems.
The
Global 200
The Global 200 is the list of ecoregions identified by the World Wide Fund for Nature ( WWF), the global conservation organization, as priorities for conservation. According to WWF, an ecoregion is defined as a "relatively large unit of land or w ...
, a set of ecoregions identified by WWF whose conservation would achieve the goal of saving a broad diversity of the Earth's ecosystems, includes a number of areas highlighted for their freshwater
biodiversity
Biodiversity or biological diversity is the variety and variability of life on Earth. Biodiversity is a measure of variation at the genetic (''genetic variability''), species (''species diversity''), and ecosystem (''ecosystem diversity'') l ...
values. The Global 200 preceded Freshwater Ecoregions of the World and incorporated information from regional freshwater ecoregional assessments that had been completed at that time.
See also
*
Crisis ecoregion
*
Lists of ecoregions
References
Bibliography
Sources related to the WWC scheme:
* Main papers:
** Abell, R., M. Thieme, C. Revenga, M. Bryer, M. Kottelat, N. Bogutskaya, B. Coad, N. Mandrak, S. Contreras-Balderas, W. Bussing, M. L. J. Stiassny, P. Skelton, G. R. Allen, P. Unmack, A. Naseka, R. Ng, N. Sindorf, J. Robertson, E. Armijo, J. Higgins, T. J. Heibel, E. Wikramanayake, D. Olson, H. L. Lopez, R. E. d. Reis, J. G. Lundberg, M. H. Sabaj Perez, and P. Petry. (2008). Freshwater ecoregions of the world: A new map of biogeographic units for freshwater biodiversity conservation. ''BioScience'' 58:403-414
** Olson, D. M., Dinerstein, E., Wikramanayake, E. D., Burgess, N. D., Powell, G. V. N., Underwood, E. C., D'Amico, J. A., Itoua, I., Strand, H. E., Morrison, J. C., Loucks, C. J., Allnutt, T. F., Ricketts, T. H., Kura, Y., Lamoreux, J. F., Wettengel, W. W., Hedao, P., Kassem, K. R. (2001). Terrestrial ecoregions of the world: a new map of life on Earth. ''Bioscience'' 51(11):933-938
** Spalding, M. D. et al. (2007). Marine ecoregions of the world: a bioregionalization of coastal and shelf areas. ''BioScience'' 57: 573–583
* Africa:
** Burgess, N., J.D. Hales, E. Underwood, and E. Dinerstein (2004). ''Terrestrial Ecoregions of Africa and Madagascar: A Conservation Assessment.'' Island Press, Washington, D.C.
** Thieme, M.L., R. Abell, M.L.J. Stiassny, P. Skelton, B. Lehner, G.G. Teugels, E. Dinerstein, A.K. Toham, N. Burgess & D. Olson. 2005. ''Freshwater ecoregions of Africa and Madagascar: A conservation assessment.'' Washington DC: WWF
* Latin America
** Dinerstein, E., Olson, D. Graham, D.J. et al. (1995). ''A Conservation Assessment of the Terrestrial Ecoregions of Latin America and the Caribbean.'' World Bank, Washington DC.
** Olson, D. M., E. Dinerstein, G. Cintron, and P. Iolster. 1996. ''A conservation assessment of mangrove ecosystems of Latin America and the Caribbean''. Final report for The Ford Foundation. World Wildlife Fund, Washington, D.C.
** Olson, D. M., B. Chernoff, G. Burgess, I. Davidson, P. Canevari, E. Dinerstein, G. Castro, V. Morisset, R. Abell, and E. Toledo. 1997. ''Freshwater biodiversity of Latin America and the Caribbean: a conservation assessment''. Draft report. World Wildlife Fund-U.S., Wetlands International, Biodiversity Support Program, and United States Agency for International Development, Washington, D.C.
* North America
** Abell, R.A. et al. (2000). ''Freshwater Ecoregions of North America: A Conservation Assessment'' Washington, DC: Island Press
** Ricketts, T.H. et al. 1999. ''Terrestrial Ecoregions of North America: A Conservation Assessment''. Washington (DC): Island Press
* Russia and Indo-Pacific
** Krever, V., Dinerstein, E., Olson, D. and Williams, L. 1994. ''Conserving Russia's Biological Diversity: an analytical framework and initial investment portfolio''. WWF, Switzerland.
** Wikramanayake, E., E. Dinerstein, C. J. Loucks, D. M. Olson, J. Morrison, J. L. Lamoreux, M. McKnight, and P. Hedao. 2002. ''Terrestrial ecoregions of the Indo-Pacific: a conservation assessment''. Island Press, Washington, DC, USA
Others:
* Brunckhorst, D. 2000. ''Bioregional planning: resource management beyond the new millennium.'' Harwood Academic Publishers: Sydney, Australia.
* Busch, D.E. and J.C. Trexler. eds. 2003. Monitoring Ecosystems: Interdisciplinary approaches for evaluating ecoregional initiatives. Island Press. 447 pages.
External links
WWF WildFinder(interactive on-line map of ecoregions with additional information about animal species)
*
Original web pageActivist network cultivating Ecoregions/Bioregions*
Original web page
{{Authority control
Ecoregions,
Biogeography